_Beggiatoa_ (parallel with the Blue-green Alga _Oscillaria_). Long
filaments formed of cylindrical cells which are attached by one
of the ends, but which are nearly always free when observed. The
filaments, like those of _Oscillaria_, describe conical figures in
their revolutions, the free filaments slide upwards and parallel with
one another; sheaths are wanting; strongly refractive sulphur drops
are found in the interior. The Beggiatoas are the most prevalent
_Sulphur-bacteria_. They occur, very commonly in large numbers,
wherever plant or animal remains are decaying in water in which
sulphuretted hydrogen is being formed; thus, for example, _B. alba_
(Fig. 31) occurs frequently as a white covering or slimy film on mud
containing organic remains. ~_B. mirabilis_ is remarkable for its
size and its strong peristaltic movements.~ The Sulphur-bacteria
oxidize the sulphuretted hydrogen, and accumulate sulphur in the
shape of small granules of soft amorphic sulphur, which in the living
cell never passes over into the crystalline state. They next oxidize
this sulphur into sulphuric acid, which is immediately rendered
neutral by absorbed salts of calcium, and is given off in the form
of a sulphate, thus CaCO_{3} is principally changed into CaSO_{4}.
In the absence of sulphur the nutritive processes are suspended, and
consequently death occurs either sooner or later. The Sulphur-bacteria
may exist and multiply in a fluid which only contains traces of
organic matter, in which organisms devoid of chlorophyll are not able
to exist. The Beggiatoas very frequently form white, bulky masses
in sulphur wells and in salt water, the traces of organic material
which the sulphur water contains proving sufficient for them. ~The
cellulose-fermentation, to which the sulphur wells in all probability
owe their origin, mainly procures them suitable conditions for
existence. The CaCO_{3} and H_{2}S, formed during the cellulose
fermentation by the reduction of CaSO_{4} is again changed into
CaSO_{4} and CO_{2} by the Sulphur-bacteria (Winogradsky, 1887).--Other
Sulphur-bacteria, the so-called purple Sulphur-bacteria, _e.g._
_B. roseo-persicina_, _Spirillum sanguineum_ (Fig. 23), _Bacterium
sulfuratum_, etc., have their protoplasm mixed with a red colouring
matter (bacterio-purpurin) which, like chlorophyll, has the power,
in the presence of light, of giving off oxygen (as proved by T. W.
Englemann, 1888, in oxygen-sensitive Bacteria). The three purple
Sulphur-bacteria mentioned, are, according to Winogradsky, not
pleomorphic kinds but embrace numerous species.~
Many _Spirilli_ (_Spirillum tenue_, _S. undula_, _S. plicatile_, and
others) are found prevalent in decaying liquids.
Bacteria (especially Bacilli) are the cause of many substances emitting
a foul odour, and of various changes in milk.
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