[Illustration: FIG. 51.--_Hydrodictyou reticulatum. A_ A cell
where the zoospores are on the point of arranging themselves to
form a net. _B_ A cell with gametes swarming out.]
The cœnobium of _Hydrodictyon reticulatum_ (Water-net) is formed of
a large number of cells which are cylindrical, and attached to one
another by the ends (Fig. 51). The asexual reproduction takes place by
zoospores, which are formed in large numbers (7,000–20,000) in each
mother-cell, within which they move about for a time, and then come
to rest and arrange themselves into a new net (Fig. 51 _A_) which is
set free by the dissolution of the wall of the mother-cell, grows,
and becomes a new cœnobium. The sexual reproduction takes place by
gamete-conjugation. The gametes are formed in the same manner as the
zoospores, but in larger numbers (30,000–100,000), and swarm out of the
mother-cell (Fig. 51 _B_). The zygote forms, on germination, 2–5 large
zoospores, each with one or two cilia, these generally swarm about
for a time, and after a period of rest become irregular thorny bodies
(polyhedra); their contents again divide into zoospores, the thorny
external coating of the polyhedra is cast off, and the zoospores,
surrounded by the dilated internal coating, unite to form a small
family, which produces several others in the manner described.
Family 2. =Confervoideæ.=
The individuals are always multicellular, the cells firmly bound
together and united into unbranched or branched filaments, expansions,
or masses of cells which grow by intercallary divisions or have
apical growth. In the first seven orders the cells are uninuclear,
but the cells of the remaining three orders contain several nuclei.
Asexual reproduction by zoospores, akinetes or aplanospores. Sexual
reproduction by isogamous or oogamous fertilisation.
The Confervoideæ, through the Ulvaceæ, are connected with the
Tetrasporaceæ, and from the _Coleochætaceæ_, which is the most
highly developed order, there are the best reasons for supposing
that the Mosses have taken their origin. The _Cladophoraceæ_
show the nearest approach to the _Siphoneæ_.
Order 1. =Ulvaceæ.= The thallus consists of one or two layers of
parenchymatous cells, connected together to form either a flat membrane
(_Monostroma_, _Ulva_) or a hollow tube (_Enteromorpha_), and may be
either simple, lobed, or branched. Reproduction takes place by detached
portions of the thallus; or asexually by zoospores or akinetes.
Gamete-conjugation is known to take place in some members of this
order, the zygote germinating without any resting-stage. The majority
are found in salt or brackish water.
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