Borelli's study is divided into a series of propositions in which he traces the principles of flight, and the mechanical actions of the wings of birds. The most interesting of these are the propositions in which he sets forth the method in which birds move their wings during flight and the manner in which the air offers resistance to the stroke of the wing. With regard to the first of these two points he says: 'When birds in repose rest on the earth their wings are folded up close against their flanks, but when wishing to start on their flight they first bend their legs and leap into the air. Whereupon the joints of their wings are straightened out to form a straight line at right angles to the lateral surface of the breast, so that the two wings, outstretched, are placed, as it were, like the arms of a cross to the body of the bird. Next, since the wings with their feathers attached form almost a plane surface, they are raised slightly above the horizontal, and with a most quick impulse beat down in a direction almost perpendicular to the wing-plane, upon the underlying air; and to so intense a beat the air, notwithstanding it to be fluid, offers resistance, partly by reason of its natural inertia, which seeks to retain it at rest, and partly because the particles of the air, compressed by the swiftness of the stroke, resist this compression by their elasticity, just like the hard ground. Hence the whole mass of the bird rebounds, making a fresh leap through the air; whence it follows that flight is simply a motion composed of successive leaps accomplished through the air. And I remark that a wing can easily beat the air in a direction almost perpendicular to its plane surface, although only a single one of the corners of the humerus bone is attached to the scapula, the whole extent of its base remaining free and loose, while the greater transverse feathers are joined to the lateral skin of the thorax. Nevertheless the wing can easily revolve about its base like unto a fan. Nor are there lacking tendon ligaments which restrain the feathers and prevent them from opening farther, in the same fashion that sheets hold in the sails of ships. No less admirable is nature's cunning in unfolding and folding the wings upwards, for she folds them not laterally, but by moving upwards edgewise the osseous parts wherein the roots of the feathers are inserted; for thus, without encountering the air's resistance the upward motion of the wing surface is made as with a sword, hence they can be uplifted with but small force. But thereafter when the wings are twisted by being drawn transversely and by the resistance of the air, they are flattened as has been declared and will be made manifest hereafter.'