Previously to the discovery last mentioned, it was by no means uncommon for paleontologists to speak with confidence of fish as having been created before reptiles. It was deemed reasonable to suppose that the introduction of a particular class or order of beings into the planet coincided, in date, with the age of the oldest rock to which the remains of that class or order happened then to have been traced back. To be consistent with themselves, the same naturalists ought now to take for granted that reptiles were called into existence before fish. This they will not do, because such a conclusion would militate against their favourite hypothesis of an ascending scale, according to which Nature "evolved the organic world," rendering it more and more perfect in the lapse of ages. In our efforts to arrive at sound theoretical views on such a question, it would seem most natural to turn to the marine invertebrate animals as to a class affording the most complete series of monuments that have come down to us, and where we can find corresponding terms of comparison, in strata of every age. If, in this more complete series of her archives, Nature had really exhibited a more simple grade of organization in fossils of the remotest antiquity, we might have suspected that there was some foundation of facts in the theory of successive development. But what do we find? In the Lower Silurian there is a full representation of the Radiata, Mollusca, and Articulata proper to the sea. The marine Fauna, indeed, in those three classes, is so rich as almost to imply a more perfect development than that which now peoples the ocean. Thus, in the great division of the Radiata, we find asteroid and helianthoid zoophytes, besides crinoid and cystidean echinoderms. In the Mollusca of the same most ancient epoch M. Barrande enumerates, in Bohemia alone, the astonishing number of 253 species of Cephalopoda. In the Articulata we have the crustaceans, represented by more than 200 species of Trilobites, not to mention other genera. It is only then, in reference to the Vertebrata, that the argument of degeneracy in proportion as we trace fossils back to older formations can be maintained; and the dogma rests mainly for its support on negative evidence, whether deduced from the entire absence of the fossil representatives of certain classes in particular rocks, or the low grade of the first few species of a class which chance has thrown in our way.