A Mechanico-Physiological Theory of Organic EvolutionNägeli, Carl
Science
A Mechanico-Physiological Theory of Organic Evolution
Nägeli, Carl
Evolution
_Law of Phylogenetic Combination._--The simplest of all plants are cells
of round form, which grow and reproduce themselves by division, budding
or free cell formation. From the fact that the younger generation of
cells, instead of separating from each other and growing to independent
plant individuals, remain united with each other, multicellular plants
arise from unicellular. The same transformation of the reproductive
cells into non-separable tissue cells is repeated several times in
multicellular plants and serves to enlarge the individual. There is
manifested in this phylogenetic process the tendency of the plant to
combine in the higher stages into one complex whole those parts which in
the lower stages tend to be independent. A similar unifying tendency is
revealed also in those plant members which have arisen by
differentiation and represent a system only by their being connected at
certain points. These combine in the higher stages and form ultimately
continuous tissues.
_Law of Phylogenetic Complication or Ampliation, Differentiation and
Reduction._--The cells, and, in general, the parts of plants which lie
near each other in space or follow upon each other in time, are always
alike in the lower stages. By differentiation they become unlike, so
that the sum of the functions which at first fall to the lot of all
parts without distinction now is shared among the individual parts. By
this means each part can perform its own special function so much the
better. Differentiation is repeated in the course of the phylogeny,
since at first all parts of an ontogeny diverge into two or more parts,
then the parts of these parts divide again, etc. Along with this process
of division another process is always active, which, as it were,
prepares the way for the former, namely, ampliation, by virtue of which
the growth of the whole ontogeny or of single stages of it undergoes a
quantitative increase, so that an organ acquires a greater number of
cells, and an individual a greater number of organs. After this increase
in number of parts in a stage of ontogeny, differentiation follows as
far as the nature of the functions permits, by the parts most separated
passing into each other by intermediate gradations. By the further
phylogenetic process of reduction the intermediate forms are suppressed.
At last only the extreme products of differentiation lie near each other
in space or follow upon each other in time; and these products are as
limited in quantity and number as possible.
Along with the above named phylogenetic processes, which take place by
the automatic increase of the idioplasm, external influences are always
active. These lend to the organism at times a local stamp corresponding
to its environment, and follow the law of adaptation.
21. ALTERNATION OF GENERATIONS IN RELATION TO PHYLOGENY.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account