A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
_Monstrous Variety._--In the British Museum, there is a dried and
somewhat injured specimen of a monstrous variety, the _Pamina
trilineata_ of J. E. Gray: it differs from the common form only in
having a tubular projection, just behind the notch separating the upper
points of the terga; this tube springs from over the terga, and is,
therefore, in a different position from the ear-like appendages in
_Conchoderma aurita_. It does not open into the sack: the membrane
composing it appears to have been double in the upper part, and to have
been lined with corium: in short, this tube seems to have been an
excrescence or tumour, of a cup or tubular form.
_General Remarks._--It will have been seen how much subject to variation
the valves of this species are. When I first examined the _Cineras
chelonophilus_ of Leach, from 36° N. lat., Atlantic Ocean, and found in
many specimens, both old and young, that the terga were very small,
flat, acuminated at both ends, with a projecting shoulder on the carinal
margin, and situated at about their own length from the apex of the
carina, and at twice their own length from the scuta; and when I found
the carina acuminated at both ends, and the scuta very imperfectly
calcified, with the lateral lobe broad, flat, and standing out at right
angles; and lastly, when I found the whole capitulum bluntly pointed,
instead of being square on the summit, I had not the least doubt, that
it was a quite distinct species. Afterwards, I found in the _Cineras
Olfersii_ of Leach, from the South Atlantic, the same form of terga; but
within slightly more concave or furrowed, and not nearly so small, and
therefore not placed at above half so great a distance from the other
valves; and here, the carina had its usual outline, as had nearly the
scutum on one side, whereas, on the other side, it presented a new and
peculiar form, having five ridges or points, and was imperfectly
calcified; seeing this, it was impossible to place much weight in the
precise form or size (and therefore, relative separation,) of the
calcified valves; and on close examination, I found every part of the
mouth and cirri identical in Leach’s _Cineras chelonophilus_ and _C.
Olfersii_, and in the common form. Therefore, I conclude, that _C.
chelonophilus_, and still more _C. Olfersii_, are only varieties; the
terga presenting the greatest, yet variable, amount of difference,
namely, in their acumination and flatness. We know, also, that in the
species of the closely allied genus of Lepas, the terga are very
variable in shape, and this is the case, even in a still more marked
degree, in _Conchoderma aurita_. Professor Macgillivray, I may add, has
come to a similar conclusion regarding the extreme variability of the
valves of this species.
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