A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
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A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
_Maxillæ_, small, narrow, produced, without any notch, with two large
upper spines, of which one is much thicker than the other; on the convex
upper margin there are some minute tufts of very small hairs.
_Outer Maxillæ_, with few bristles, arranged in a continuous line on the
anterior surface; on the external surface there is a tuft of long
bristles. Olfactory orifices situated laterally, forming two flattened,
tubular projections.
_Cirri._--First pair placed not far from the second; the three posterior
pair not very long, with their segments elongated, not protuberant,
bearing four pair of non-serrated spines, with a single short bristle
between each pair; dorsal tufts small, with one spine longer than the
others. First cirrus rather short, segments not very broad; second
cirrus with the rami nearly equal in length, anterior ramus rather
thicker than the posterior ramus, with three longitudinal rows of
spines.
_Caudal Appendages._--These are minute, rather broad, not half as long
as the lower segments of the pedicels of the sixth cirrus, with four
very long spines at the tip.
_Penis._--There is no trace of a probosciformed penis in the four
specimens examined; and as this organ is present in every ordinary
cirripede, with the exception of _Ibla Cumingii_ which we know to be
exclusively female, so we may infer with some confidence that the form
here described is female, although it is impossible in specimens once
dried to demonstrate the absence of the vesiculæ seminales and testes.
_Affinities._--This is a very distinct species; it is, however, much
more nearly related to _S. rutilum_, than to any other species; and next
to this, to _S. vulgare_; from this latter species it chiefly differs in
the large scales of the peduncle, in the scuta not being added to at
their upper ends, and in the membrane covering and connecting the valves
being spineless; but there is a greater difference in the trophi and in
the cirri. The peduncle of _S. ornatum_ presents some resemblance to
that of the singular cretaceous genus, _Loricula_.
MALE.
All the specimens, as already stated, were dry, but in an excellent
state of preservation, so that after having been soaked in spirits, they
could be minutely examined. In the four which I opened, I found, in a
transverse pouch on the under side of each scutum, a male lodged; in a
fifth dead and bleached specimen, the cavities in the shell for the
reception of the males, were present; and in a sixth young specimen,
also dead, cavities were in process of formation. As compared with
plants, the relation of the sexes in this species may be briefly given,
by saying that it belongs to the class _Diandria monogynia_. I will
first describe the males themselves, and then the cavities in the shell
of the female. The males differ in every point of detail, from the
complemental males of _S. vulgare_, but yet present so close a general
resemblance, that a comparative description will be most convenient.
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