A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
carina they are trifid and quadrifid, and large enough to be conspicuous
with a lens of weak power.
_Peduncle._--The most remarkable fact concerning this part, is that the
outer tunic, together with the calcareous scales with which it is
covered, is moulted at each successive period of exuviation and growth.
I demonstrated this fact in _L. dorsalis_ and _L. truncata_, by removing
the old tunic and finding a new membrane with perfect calcified scales
beneath; and as these two species, (I obtained, also, pretty good
evidence in _L. Nicobarica_,) are at the opposite extremes of the genus,
no doubt this fact is common to the whole genus. I know of no other
instance, amongst Cirripedia, in which _calcified_ valves or scales are
moulted. I am not certain that the whole skin of the peduncle is thrown
off in a single piece; though this almost certainly is the case with the
uppermost and lowest portions. The animal’s body is partly lodged within
the peduncle, which is generally from one to three times as long as the
capitulum, and in the upper part is fully as broad as it. The scales
with which it is clothed, extend up in the triangular interspaces
between the basal margins of the valves. The scales of the upper whorl,
or of the two or three upper whorls (Pl. VIII, figs. 1 _b´_ and 3 _d_)
are larger than those below; and these latter rapidly decrease in size,
so as to become low down on the peduncle, almost or quite invisible to
the naked eye. The scales in each whorl, are placed alternately with
those in the whorls, above and below. All the upper scales are packed
rather closely together; those in the uppermost row are generally nearly
quadrilateral; those in the few next succeeding whorls, are triangular,
with their basal margins protuberant and arched; the scales, low down on
the peduncle, stand some way apart from each other, and generally
consist of simple rounded calcareous beads, of which some of the
smallest in _L. dorsalis_ were only 1/400th of an inch in diameter. In
the lowest part of the peduncle these scales, after each fresh
exuviation, are apparently soon worn entirely away by the friction
against the sides of the cavity; hence in most specimens this part of
the peduncle is quite naked. This same part, however, is furnished with
nail- or rather star-headed little projections of hard, yellow, horny
chitine (fig. 3 _e_). The star on the summit seems generally to have
about five irregular points; one star which I measured was 7/6000th of
an inch in total width, the footstalk being only 2/6000th of an inch in
diameter; the whole projected 10/6000ths of an inch above the surface of
the peduncle; from the footstalk a fine tubulus runs through the
membrane to the underlying corium. These star-headed little points are
often much worn down; in one specimen which was on the point of
exuviation, there remained, in the lower part, close above the basal
calcareous cup, only some hard, smooth, yellow, little discs, on a level
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