A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
with the general surface of the membrane,--these being the intersected
or worn down footstalks, with every trace of the calcareous beads gone.
But in this same specimen, under the old peduncular membrane, there was
a new one, studded with the usual circular calcareous beads, slightly
unequal in size, generally about 1/400th of an inch in diameter, and
each furnished with a tubulus; but as yet none of the star-headed points
of chitine had been formed. I believe that these latter are developed
from the tubuli leading to the calcified beads, and, therefore, are
formed directly under them. In _L. cauta_ the lowest scales on the
peduncle are a little larger than in _L. dorsalis_, giving a frosted
appearance to it, and all of them are serrated (fig. 3 _d_) round their
entire margins. Generally only the scales in the uppermost, or in the
three or four upper rows are serrated, and this only on their arched and
protuberant lower margins. The state of the serrated edge varies
extremely in the same species, from elongated conical teeth to mere
notches, according to the amount of wear and tear the individual has
suffered since the last period of exuviation; so also do the teeth or
serrated margins on the valves of the capitulum. Each scale has a fine
tubulus passing from the corium through the membrane of the peduncle to
its bluntly-pointed imbedded fang or base. The membrane is transparent,
thin, and tender, to a degree I have not seen equalled in the other
Lepadidæ, except, perhaps, in Ibla. It is much wrinkled transversely.
_Muscles of the Peduncle._--These consist of the usual interior and
longitudinal,--exterior and transverse--and oblique fasciæ; the former
are unusually strong; downwards they are attached to the basal
calcareous cup or disc, and upwards they extend all round to the lower
curved margins of the valves. They are, as usual, without transverse
striæ. Besides these, there are, (at least in _L. dorsalis_ and _L.
Nicobarica_,) two little fans of striæ-less muscles, which occur in no
other pedunculated cirripede; they are attached on each side of the
central line of the carina, near its base; they extend transversely and
a little upwards, and each fan converges to a point where the lower
margins of the carina and terga touch; of these muscles, the upper
fasciæ are the longest. Their action, I conceive, must be either to draw
slightly together the basal points of the terga, and so serve to open
their occludent margins, or to draw inwards the base of the carina:
these muscles apparently first shadow forth the posterior or carinal,
transversely-striated, opercular muscles of sessile cirripedes.
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