A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes — John Shaqi
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
serve as a lower lip; they are thicker than the other trophi; they have
their inner surfaces clothed with spines, sometimes divided into an
upper and lower group, and occasionally separated by a deep notch: there
are often long bristles outside. They are furnished with at least two
muscles; in sessile Cirripedes I have seen that they are capable of a
rapid to and fro movement, and I have no doubt that their function is to
brush any small creature, caught by the cirri, towards the maxillæ,
which are well adapted to aid in securing the prey, and to hand it over
to the mandibles, by them to be forced down the oesophagus. On the
exterior face of the outer maxillæ, above a trace of an upper
articulation, either two small orifices or two large tubular projections
can always be discovered; and these, as will presently be mentioned, I
believe to be olfactory organs.
_Cirri._--The five posterior pair are seated close to each other and
equidistant; the first pair is generally seated at a little distance,
and sometimes at a considerable distance from the second pair. The first
pair is the shortest; the others, proceeding backwards, increase
gradually in length. The rami of each pair are either equal in length or
slightly unequal: those of the first pair are oftenest unequal. The
number of segments in the posterior cirri is sometimes very great; in
one species of Alepas, there were above sixty segments in one ramus, the
other ramus being in this unique case (Pl. X, fig. 28) small and
rudimentary. The pedicels consist of two segments, a lower, longer, and
upper short one (fig. 18, _c_, _d_.) In the usual arrangement of the
spines on the segments of the three posterior pair of cirri, there are
(figs. 26, 27) from three to six pair of long spines on the anterior
face, with generally some minute spines (occasionally forming a tuft)
intermediate between them: on the dorsal surface, in the uppermost part
of each segment, there is a tuft of short spines generally mingled with
some longer, finer ones: on the inner side of each segment, on the upper
rim, there are generally a few extremely minute and short spines. From
the increase of these latter and of the intermediate spines, the
antero-lateral faces of the segments of the first cirrus, and of the
lower segments of the anterior ramus of the second cirrus (Pl. X, fig.
25), are almost always thickly paved with brush-like masses of spines.
The lower segments of the anterior ramus of the third cirrus is
generally, though not always, thus paved: these paved segments are much
broader than the others. The posterior rami of the second and third
cirri are often in some slight degree paved, though in other cases they
resemble the three posterior pair of cirri. The two segments of the
pedicels have bristles on their anterior faces, essentially arranged on
the same plan as on the segments of the rami: the bristles are generally
not so symmetrically arranged on the pedicels of the second and third
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