A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes — John Shaqi
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
a distinct organ, like the closed tube in certain Annelids.
Occasionally, I have seen one imperfect epithelial bag or tube within
another and later-formed one. Digestion seems to go on at the same rate
throughout the whole length of the stomach; if there be any difference,
the least digested portions lie in the lower and narrower part. The
prey, consisting generally of crustacea, infusoria, minute spiral
univalves, and often of the larvæ of Cirripedes, is not triturated: when
the nutritious juices have been absorbed, the rejectamenta are cast out
through the anus, all kept together in the epithelial bag, which is
excluded like a model of the whole stomach, with the exception of that
part coated by the bell of the oesophagus. I have sometimes thought that
the bag was formed so strong, for the sake of thus carrying out the
excrement entire, so as not to befoul the sack. I believe Lepas can
throw up food by its oesophagus; at least, I found in one case, many
_half-digested_ small Crustaceans in the sack, and others of the same
kind in the stomach.
_Circulatory System._--I can add hardly anything to what little has been
given by M. Martin St. Ange: like others, I have failed, as yet, in
discovering a heart. The whole body is permeated by channels, which have
not any proper coat: there is one main channel along the ventral surface
of the thorax, dividing and surrounding the mouth, and giving out
branches which enter the inner of the two channels in each cirrus: as
Burmeister has shown, there are also two channels in the penis. There
are two dorso-lateral channels in the prosoma, which are in direct
connection with the great main channel, running down the rostral (_i.
e._, ventral) side of the peduncle. This latter main channel branches
out in the lower part, and transmits the fluid through the ovarian
tubes, whence, I believe, it flows upwards and round the sack,
re-entering the body near the sides of the adductor scutorum muscle. The
main rostral channel (or artery?) in the uppermost part of the peduncle,
has a depending curtain, which, I think, must act as a valve, so as to
prevent the circulating fluid regurgitating into the animal’s body
during the contractions of the peduncle.
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