A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
Sessile Cirripedes, partly from being attached to surfaces having
very different characters, partly from undergoing a varying amount of
disintegration, and partly from unknown innate causes, are extremely
variable. Under the head of _Variation_, in the Family Balanidæ and
under the Genus Balanus, I have enlarged on this subject, and have
shown that there is scarcely a single _external_ character which is not
highly variable in most of the species. As whole groups of specimens
often vary in exactly the same manner, it is not easy to exaggerate the
difficulty of discriminating species and varieties. It is absolutely
necessary in most cases, for mere identification, that the valves of at
least one specimen in a group should be disarticulated and well cleaned
(for which purpose caustic potash is most useful), so that the internal
characters may be examined. Whoever attempts to make out from external
characters alone, without disarticulating the valves, the species,
(even those inhabiting one very confined region, for instance the
shores of Great Britain,) will almost certainly fall into many errors:
hence it is, and can thus only be accounted for, that I have not seen
one collection of British specimens with _all_ the species, though so
few in number, rightly discriminated; and in the large majority of
cases, either two or three species, certainly distinct, were confounded
together, or two or three varieties, as certainly not distinct, were
separated from each other.
_On the Names Given to the Different Parts of Cirripedes._
In my former volume I have stated that I found it indispensable, in
part owing to the extreme confusion of the nomenclature previously
used, to attach new names to several of the external parts of
Cirripedes. Almost all these names are applicable to the Balanidæ,
or sessile Cirripedes, and to the Verrucidæ; but a few additional
names are requisite, which, together with the old names, will, I
hope, be rendered clear by the accompanying woodcuts. In sessile
Cirripedes, the whole of that which is externally visible, may for
convenience sake be divided into the _operculum_ or _opercular valves_
(_valvæ operculares_), and the _shell_ (_testa_), though these parts
homologically present no real difference. The operculum is seated
generally some little way down within the _orifice_ of the shell; but
in very young specimens and in Verruca, the operculum is attached
to the summit of the shell, and the shell, without the operculum being
removed, can hardly be said to have any orifice; though, of course, the
opercular valves themselves have an aperture for the protrusion of the
cirri.
The shell consists of the _basis_ (called the support by some authors),
which is membranous or shelly, and flat or cup-formed, and of the
_compartments_ (_testæ valvæ_), which vary from eight to four in
number, and occasionally are all calcified together.
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