A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
The probosciformed penis lies adpressed on the under side of the
thorax, with its apex generally projecting between the first and second
pairs of cirri. It presents the same ringed or articulated structure as
in the Lepadidæ: it arises from an unarticulated projection or support,
which also forms the posterior border to the anus. This support
often terminates, as first observed by Poli, in a very sharp point;
but this point cannot be of much functional importance, for though
present in _Balanus balanoides_, it is absent in the closely allied
_B. crenatus_; in Tubicinella there is only a rudiment of this point;
I have not observed it in any member of the Chthamalinæ. The strong,
transverse and longitudinal muscles with which the penis is furnished,
are attached to this support. The apex or orifice of the penis is, I
believe, invariably surrounded by some bristles. Its length varies
much, according to its state of contraction or relaxation; and this
again, I believe, is dependent on the condition of the male secreting
organs. In a small specimen of _Elminius modestus_, the penis was
actually thrice as long as the whole thorax, including the prosoma:
in Pachylasma and in _Octomeris angulosa_, the penis is very short,
being equal only to once and a half the length of the pedicel of the
sixth cirrus: in _Octomeris brunnea_, the unarticulated support is
much elongated, being as long as the pedicel of the sixth cirrus, in
which respect this organ resembles that of _Ibla quadrivalvis_, and of
no other Cirripede. From the attachment of the penis at the posterior
end and on the under side of the anus--from the position of the caudal
appendages (where such occur) over the anus--from the position of
these same appendages in the pupa--and lastly, from the position of
the papilla-like penis in the abnormal _Proteolepas_, I infer that,
homologically, the penis is situated at the apex of the abdomen, on
its ventral surface; and that, consequently, this organ cannot be
considered as the abdomen itself in a modified condition.
_Female Organs of Generation._
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