A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
[71] In the case of one _young_ shell, I found that the
previously-formed circumferential slip must have split, long before
it had assumed its proper and ordinary width; for the last-formed
slip of basal membrane was of extreme narrowness, and would have
to be considerably added to in width, whilst the new and narrow
circumferential slip was likewise being added to in width. This
slip of basal membrane, though so extremely narrow, had its own
cement-ducts and glands.
The central slip or rather disc of membrane, is 3/400ths of an inch in
diameter; and this shows the basal diameter of the shell immediately
after the metamorphosis. In the middle of this little disc I saw,
in several specimens, the prehensile, pupal antennæ; I made out
distinctly the ultimate segment with its bristles, and, as I believe,
the disc-segment, which was 7/2000ths of an inch in diameter; but this
portion was much obscured by the quantity of cement. When the corium
is removed from the inner side of the basal membrane, the two chains
of glands, extending from exactly over the antennæ in the centre about
half way towards the circumference, are conspicuous. The cement-trunk,
connecting the glands, is thin, and at the further end is always broken
off, by the removal of the corium and overlying layer of ovarian cæca,
into which the two cement-trunks enter; and without which removal,
nothing could be seen. The two chains of glands form a very large
angle, open towards the rostral end of the shell, as represented at
fig. 1 _c_, much enlarged; by a mistake in fig. 1 _a_, the two have
been drawn in a straight line. The cement-trunk increases in diameter
in proceeding from the centre to the circumference, and the glands
likewise increase in size, at the same time altering somewhat in shape.
From near the lower side (the basal membrane being viewed from within)
of each gland, two cement-ducts proceed, which pour out their contents
beneath the basal membrane. The orifices of the ducts always exactly
face the middle folds of the two lateral, and two carino-lateral
compartments. In a full-sized specimen, there are from thirty-five to
forty cement-glands on each side, always corresponding exactly with
the number of slips of basal membrane, including the circumferential
slip, to which the last-formed pair of glands and cement-ducts belong.
In correspondence with the great number and narrowness of the central
slips of membrane, so are the cement-glands towards the centre numerous
and very small. All the glands, in the more central parts, consist of
a mere transverse enlargement of the cement-trunk; but the exterior
and larger glands, which are more closely packed together, are more
globular or pear-shaped; and the two ducts (of which the one on the
rostral side is considerably enlarged at its base) do not come out of
the gland exactly at the same level. The trunk, connecting the glands,
runs straight from one to the other. The ducts proceeding from the
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