A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
outer and older glands, on the carinal side, are much curved (Pl. 28,
fig. 1 _c_). To give an idea of the dimensions of the several parts, I
may state that the largest ducts were 3/1000ths of an inch in diameter,
and the glands belonging to them nearly thrice as much, measured in the
direction of the cement-trunk; on the other hand, some of the ducts
from the small central glands had a diameter eighteen times less than
that of the largest ducts. Towards the circumference, the ducts that
proceed from the older and larger glands are piled one exactly over the
other--the last formed being the topmost, and all are imbedded in the
corium which overlies the basal membrane: this position of the ducts,
one over the other (which could not be well represented in figs. 1 _a_
and 1 _c_), is owing to their all debouching at the same exact point.
But the ducts form the smaller and younger glands, when the shell
had a different shape, are spread out, and are all attached to the
basal membrane. Altogether, the basal membrane of Coronula, when well
cleaned, and examined under a moderately high power, presents the most
singular and elegant appearance.
We now come to the cement-tissue: this lies on the under or outer
side of the basal membrane; it is not represented in figs. 1 _a_ or
1 _c_, but only in the sectional diagram, 1 _b_, by the letters _z_,
_z_: it presents its usual character and appearance, like solid glue
or brown gum, but is obliquely laminated and sub-laminated: it forms
a layer, much thicker than the basal membrane itself, being as much
as .004 or .005 of an inch in thickness. It is, however, difficult
to ascertain its thickness, from the singular manner in which it
penetrates into and almost blends with the epidermis of the whale's
skin; so much so, that for a considerable time I thought (not then
knowing anything about the cement of Cirripedes) that this transparent
horny substance probably answered to a corn on the human foot produced
by pressure. But I soon observed that this horny substance certainly
extended into and up the cement-ducts; and this fact first led me to
the examination of the whole subject in the several genera of Lepadidæ
and Balanidæ. It was not difficult to remove the cement-ducts, leaving
small portions of the contained cement projecting freely up as points
from the general surface of cement. The cement adheres slightly to the
whole basal membrane, but chiefly to the yellowish rims or edges of
the successive slips; and it is indeed this adhesion which, I believe,
produces the rims; for the circumferential slip, when first split, had
very thin ragged edges. The cement also extends under the spoke-like
prolongations of the circumferential slip, and likewise some way up the
sides of the walls.
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