A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
The cement-glands, the trunk, and the ducts, except the two, three,
or even four last-formed ones, are all filled with an apparently
solid thread of transparent, brownish cement, differing in no respect
from the cement under the central parts of the basal membrane. In
one instance, in which the last-formed pair of glands and ducts had
apparently been only just developed, they were so perfectly transparent
that I could distinguish them only under certain lights, and I could
not perceive any contents. The last-formed glands and ducts always
appear very delicate, and include a tube of very delicate tissue,
containing more or less of granular matter. The next succeeding pair of
glands and ducts are always more opaque, and contain much more granular
matter; which, in the next, or next but one, may be seen passing into
the state of pale brown, transparent chitine. I have seen some most
distinct instances, in which, in the same duct, the part towards the
centre of the basis was filled with homogeneous cement, and the part
towards the circumference with still plainly granular matter. In the
successive circular slips of cement-tissue, lying attached under the
circumferential slip and under the two or three succeeding slips of
basal membrane, an exactly analogous series of changes is presented; as
indeed might have been expected, as the slips of cement are absolutely
continuous with the contents of the ducts. Moreover, if a vertical
section be made across one of the last-formed slips of cement, it
may sometimes be seen to be apparently in the act of separating into
layers, with the lower layers more pulpy, elastic, and white than the
upper layers, which are less coherent, and show as yet even still less
the character of cement. The cement under the circumferential and
adjoining slips, often presents a peculiar wrinkled appearance, in
lines conformable with the outline of the basal membrane; but I do not
believe that these are real wrinkles, though so like them; they seem
to consist of sinuous threads, longer or shorter, sometimes slightly
branched, crossing and inter-joined, and composed apparently of faintly
coloured cement. I suspect that these threads are formed by the union
and subsequent drawing out of aggregations of that matter, which within
the ducts is first granular, and then changes into cement; for at the
orifices of the ducts these wrinkled threads sweep outwards in curved
lines on both sides. The cement in these early stages adheres, with
very little force, to the basal membrane; and with only a little more
force to the underlying layers of cement; in fact, till it assumes the
brown translucent appearance, like solid glue, it hardly seems to act
as cement.
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