A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc. — John Shaqi
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
In the larva in the first stage, an eye and two pairs of antennæ are
in process of formation or are developed; here, then, according to
the analogy of all Crustaceans, we have evidence of the existence of
the first three cephalic segments. The mouth always consists of three
pairs of gnathites, and hence again, from analogy, this part may be
inferred to be formed of, and supported on, three other segments;
making thus far six segments. In two Orders out of the three into
which Cirripedes may be divided, namely, in the Abdominalia and Apoda,
eight quite distinct segments succeed the mouth; of these the first
differs slightly from the seven succeeding segments, and may, I think,
be safely considered as forming the seventh (cephalic) segment. The
next seven segments resemble each other in all essential respects, and
are no doubt the normal, seven thoracic segments. These, in both the
above orders, are succeeded by three smaller segments, which differ
in structure from the thoracic segments, and must be abdominal. Hence
we here have, altogether, seventeen segments. It should, however, be
observed that in the two orders just referred to, each includes only
a single species; but I know of no good reason why, on this account,
their structure should be valued the less. In the third order, the
Thoracica, which includes all common Cirripedes, two segments with
their appendages are missing out of the eight that should succeed
the mouth; from the open interval in the pupa, between the mouth and
first pair of natatory legs, and from some other reasons, I believe
that the two missing segments are the seventh and eighth, or last
cephalic and first thoracic segments, and that they have coalesced
close posteriorly to the mouth.[7] In the order Thoracica, the abdomen
is quite rudimentary, though often still bearing caudal appendages;
in the pupa, however, of this order, as in the mature animal of the
two other orders, it is formed of three segments. Hence I conclude
that, notwithstanding the absence of the above two segments with their
appendages in the Thoracica, the archetype Cirripede may be safely said
to be composed of seventeen segments.
[7] This question and the whole subject of the homologies of the
several parts of a Cirripede, will be discussed under the head of
the Metamorphoses of the Balanidæ.
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