A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
_Basis._--In typical species the basis is calcareous, and consists of
an upper and lower lamina, separated by radiating septa, forming pores.
In the same manner as the septa of the parietes sometimes, though
rarely, become irregularly divided near the outer lamina, forming
outer pores, so it is, but in a much more marked degree, with the
basis. The basis in such cases becomes extremely thick, and consists
of an upper, thin lamina, with the regular radiating septa and pores,
and of an underlying, thick, cancellated mass, which seems wholly to
result from the dividing and sub-dividing of the septa. The basal
radiating pores, like the parietal pores, are closed at intervals by
calcareous transverse septa. The basal points of the parietal septa
enter the orifices of the basal pores, and the threads of corium pass
into the latter, between the denticuli of the parietal septa. In some
species, as in _B. crenatus_ and _Hameri_, the basis is perfectly
solid, the upper lamina being absent, just as in some species, the
internal lamina of the parietes is absent. In _B. flosculus_ the basis
is calcareous, but consists of so excessively thin a film as hardly
to be distinguished: it presents, moreover, as also is the case with
_B. imperator_, a beaded structure. Again, in some few species, as
in _B. balanoides_, the basis is simply membranous. When the basis
is thin, it is always flat, and is closely moulded to the irregular
surface of attachment; and in this case, when specimens are crowded
together, their elongation is effected exclusively by the growth of the
walls; but, when the basis is thick, it sometimes becomes, in crowded
groups, deeply, but irregularly, cup-formed, or cylindrical, as in _B.
psittacus_ and _perforatus_. In _B. allium_, however, which inhabits
massive corals, the basis is as regularly concave or cup-formed as
in the genus Pyrgoma: in _B. calceolus_ and its allies, and in some
varieties of the fossil _B. inclusus_, the basis is boat-formed, with
its lower surface deeply grooved longitudinally from clasping the
stem of the Gorgonia or other zoophyte, to which it was attached.
In certain varieties of _B. lævis_ it is very remarkable that the
deeply cup-formed basis becomes, owing apparently to the whole shell
having grown too deep for the animal, half-filled up with irregular,
calcareous, transverse plates (Pl. 4, fig. 2 _a_), resting one upon the
other by irregular points or pillars. The cementing apparatus has been
sufficiently described under the Family.
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