A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
_Mouth._--The _labrum_ is always notched; sometimes it has no teeth,
but generally there are three on each side; in _B. balanoides_ there
are five or six on each side; and in _B. improvisus_ and _eburneus_
there is a whole row of teeth (Pl. 26, fig. 2, _e′_), graduated in
size, on each side of the notch. The _palpi_ are large, with their
apices nearly touching, and furnished with long spines. The _mandibles_
have, as it appears, normally, five teeth, but the two lower teeth
are always small and often rudimentary, and almost confluent with
the inferior, sometimes spinose angle. The _maxillæ_ have either a
simple edge, or a notch under the pair of large upper spines, or the
lower part forms (Pl. 26, fig. 7) a step-formed projection: there are
generally two lower spines, placed singly or not in pairs, larger than
the others, with the exception of the uppermost pair. The _outer
maxillæ_ are, on their inner faces, obscurely divided into two lobes.
_Cirri._--The rami of the first pair are unequal, the shorter one
sometimes not being more than half the length of the other ramus:
the segments of the shorter ramus are broad, and are, together with
the lower segments of the longer ramus, thickly clothed with spines;
in some species, as in _B. perforatus_, the anterior surfaces of the
segments, more especially of the shorter ramus, and of both rami of the
second pair are produced (Pl. 29, fig. 4), so as sometimes to form very
remarkable projections. The segments of the second and third pairs are
always thickly clothed with spines, as also are their pedicels. The
third pair is rather longer than the second; but in _B. vestitus_ and
_imperator_ it is much longer, and is otherwise somewhat different.
The dorsal and basal margin of the pedicel of the third pair, in some
of the species, as in _B. tintinnabulum_, is produced backwards on the
thorax, and forms a membranous plate fringed with fine spines. The
three posterior and longer pairs of cirri have from three to rarely
eight or ten pairs of long spines on each segment, with generally one
or two minute spines in the middle between each pair: their pedicels
have a regular double row of spines.
The _penis_ is long and hairy: in most of the species there is, at its
dorsal basis, a small, sharp, flattened, imperforate projection; first
observed by Poli: but this is sometimes absent, as in _B. crenatus_,
though present in the closely allied _B. balanoides_; and its presence
is variable in _B. tintinnabulum_. All the species have large plicated
_branchiæ_. The base of the _sack_ in several species is furnished
with inwardly projecting filamentary appendages. In _B. perforatus_,
_crenatus_, and _improvisus_, and I believe in other species, the upper
part of the _stomach_ is furnished with a circle of branching cæca.
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