A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
_On the variation of the species_; _their arrangement and affinities_;
_value of the characters used_; _changes during growth_.--Owing to the
great variation in external characters, to which almost all the species
are subject, and likewise to the genus being a very natural one, that
is, to the species following each other in close and natural order, it
is not easy to exaggerate the difficulty of identifying the species,
except by a deliberate examination of the internal and external
structure of each individual specimen. Every one who has collected
sessile cirripedes must have perceived to what an extent their shape
depends on their position and grouping. The surface of attachment has
a great effect on that of the shell; for as the walls are added to at
their bases, every portion has at one time been in close contact with
the supporting surface; thus I have seen a strongly-ribbed species
(_B. porcatus_) and a nearly smooth species (_B. crenatus_) closely
resembling each other, and both having a peculiar appearance, owing
to their having been attached to a pecten. Dr. Gray has pointed out
to me specimens of _B. patellaris_, curiously pitted like the wood
to which they had adhered; and numberless other instances might be
added. Quite independently of the effect produced by the surface of
attachment, the degree to which the longitudinal folds and ribs are
developed on the parietes, is variable in most of the species, as in
_B. tintinnabulum_, _vestitus_, and even in _B. porcatus_; the presence
or entire absence of these ribs often surprisingly alters the whole
aspect of the shell. The persistence of the so-called epidermis is in
some degree variable; and in _B. lævis_ we have groups of specimens
absolutely naked, and others uniformly clothed with a brown membrane.
Again, some species in certain localities are all subject to the
disintegration of the entire outer lamina of the walls; and in such
cases (as with _B. perforatus_) there is not the smallest resemblance
between the corroded and perfect specimens. The size of the orifice,
and consequently of the operculum, compared with the shell itself,
varies accordingly as the shell is more or less conical or cylindrical;
in the latter case, the summits of the radii are generally more oblique
and the aperture consequently more deeply toothed than in the more
conical varieties. Size is a serviceable character in some cases, but
very many specimens are required to ascertain the average or maximum
size of each species, for there is no method of distinguishing a
half-grown from a full-grown specimen; and I believe, as long as the
individual lives, so long does it go on moulting and growing. _Colour_
is of very considerable service; though the precise tint varies greatly
in almost every species; and what is a far more serious evil, the
majority of the species have their white or nearly white varieties, the
latter sometimes as numerous as the coloured ones: in _B. perforatus_,
Public-domain text, read in full here on John Shaqi.
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