A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
There are always six pairs; each biramous and multiarticulated,
supported on a pedicel formed of two segments. A shield-like swelling
at the exterior bases of these pedicels often appears like another
segment; but such, I believe, is not its nature. The five posterior
pairs answer to the five pairs of ambulatory legs in the higher
Crustaceans; and as in the case of the latter, the three, or the four
hindermost pairs almost invariably resemble each other. The first
pair, which is homologous with the outer maxillipeds of ordinary
Crustaceans, is separated by an interval from the second pair;--though
this is not the case with the legs of the pupa, from which the cirri
are metamorphosed. These anterior cirri are attached to the lateral
edges of the mouth, namely to the thickened rim of membrane, forming
the supposed basal segment of the mandibles. They are capable of more
diversified movements than the other cirri: the anterior ramus is
always elongated, with the terminal segments more or less tapering,
and is directed beyond (or anteriorly to) the mouth: the shorter
ramus closely resembles in structure the rami of the second pair.
In the Chthamalinæ the second pair, and in the Balaninæ the second
and third pairs (as will be more particularly described under these
sub-families) differ in structure from the posterior pairs, from which
they are separated by a slight interval. The number of segments on
the posterior cirri is often great, amounting in Chelonobia even to
fifty. Each segment normally is furnished on its inner face, which
is usually somewhat protuberant, with from two to rarely eight or
ten pairs of long spines or bristles, placed in a double row; the
two spines in the lower pairs stand nearer to each other, and are
shorter than the spines in the upper pairs. Between each pair of
spines there is either a single, very thin bristle, or often a tuft of
such. The pairs are directed somewhat upwards, and they diverge when
the cirri are uncurled; their function is obviously to entangle the
prey. On the dorsal or exterior surface of each segment, close to its
upper margin, there is a tuft of spines, often composed of thicker
and thinner spines; these, I believe, serve to prevent any creature
intruding within the sack. On both sides of the upper margin of each
segment, there is generally a row of short, blunt, excessively minute
spines, which only deserve notice, inasmuch as it is by their increase
in number and size, and by the spreading out of the dorsal tufts,
and, lastly, by the increase of the little tuft intermediate between
the pairs of spines situated in front, that the segments on the two
or three anterior pairs of cirri become covered, like brushes, with
bristles. The bristles or spines on the second and third cirri are
often, especially in Tetraclita, doubly and coarsely pectinated. The
bristles on the pedicels follow the same arrangement as on the rami;
namely, being in regular pairs on the posterior cirri, and crowded
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