A New Order of Fishlike Amphibia From the Pennsylvanian of Kansas — John Shaqi
A New Order of Fishlike Amphibia From the Pennsylvanian of KansasEaton, Theodore H. (Theodore Hildreth)
Science
A New Order of Fishlike Amphibia From the Pennsylvanian of Kansas
Eaton, Theodore H. (Theodore Hildreth)
Amphibians, Fossil; Paleontology -- Pennsylvanian
Probably associated with the characters of the order, as given above,
are the connection of pectoral girdle with skull, and the presence of a
tympanic membrane, the stapes functioning in both sound-transmission
and palatoquadrate suspension.
Family HESPEROHERPETONIDAE
Orbits and foramen magnum unusually large in correlation
with reduced size of animal; squamosal forming posterior
margin of orbit; circumorbital series absent (except for
postorbital); sensory pits on squamosal and frontal.
Characters defining the family are evidently the more specialized
cranial features, which probably evolved during Mississippian and early
Pennsylvanian times.
The definition of the genus and species may be left to rest upon
Peabody's (1958) original description and the present account, until
the discovery of other members of the family gives reason for making
further distinctions.
SUMMARY
_Hesperoherpeton garnettense_ Peabody (1958), based on a
scapulocoracoid and part of a vertebra, was originally placed in the
order Anthracosauria, suborder Embolomeri, family Cricotidae. A new
skeleton from the type locality near Garnett, Kansas (Rock Lake shale,
Stanton formation, Upper Pennsylvanian), shows that the animal has the
following rhipidistian characters: Large notochordal canal below
foramen magnum, otico-occipital block separate from ethmosphenoid,
postaxial processes on three axial bones of forelimb, pectoral girdle
(probably) articulated with tabular. Nevertheless, _Hesperoherpeton_
has short digits, an anthracosaurian type of pectoral girdle, an otic
rather than spiracular notch, nostrils separate from the mouth, and
vertebrae in which the intercentrum is U-shaped and the pleurocentra
large but paired. The stapes reaches the quadrate.
_Hesperoherpeton_ is placed in a new order, PLESIOPODA, on the basis of
the characters stated above, and a new family, HESPEROHERPETONIDAE.
Specialized characters of the family include: Reduction of
circumorbital bones, bringing the squamosal to the edge of the orbit,
loss of certain bones of the temporal region, and relative enlargement
of the orbits and foramen magnum, in correlation with the diminutive
size of the animal. The structural characters of _Hesperoherpeton_
suggest to us that it lived in the shallow, weedy margins of lagoons,
rested with its head partly out of water, and normally did not walk on
land.
LITERATURE CITED
EATON, T. H., JR.
1951. Origin of tetrapod limbs. Amer. Midl. Nat., 46: 245-251.
JARVIK, E.
1952. On the fish-like tail in the ichthyostegid stegocephalians.
Meddel. om Grønland, 114: 1-90.
1954. On the visceral skeleton in _Eusthenopteron_ with a discussion
of the parasphenoid and palatoquadrate in fishes. Kgl. Svenska
Vetenskapsakad. Handl., 5: 1-104.
1955. The oldest tetrapods and their forerunners. Sci. Monthly,
80: 141-154.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account