A Synopsis of Neotropical Hylid Frogs, Genus OsteocephalusDuellman, William Edward
Science
A Synopsis of Neotropical Hylid Frogs, Genus Osteocephalus
Duellman, William Edward
Frogs -- Ecuador; Osteocephalus
Cochran and Goin (1970) were unaware of the identities of _Hyla
verrucigera_ and _riopastazae_; they used the later name _Osteocephalus
orcesi_ for Colombian frogs that are correctly referred to _O.
verrucigerus_. Although Goin (1961) placed _Hyla buckleyi_ and _H.
pearsoni_ in _Osteocephalus_, Cochran and Goin (1970) recognized a
"_buckleyi_ group" in _Hyla_ that included these two species plus a new
species, _Hyla cabrerai_ from Amazonian Colombia and Brasil (total of
three specimens). Also, these authors named _Hyla carri_ from a single
Colombian specimen. Study of the types of _Hyla cabrerai_, _H. carri_,
and _H. festae_ Peracca, 1904, from Ecuador, reveal that all of these
names are synonyms of _Osteocephalus buckleyi_.
Much of the taxonomic confusion and multiplicity of trivial names is due
to the great amount of color variation in _taurinus_ and to the sexual
dimorphism in the texture of the dorsal skin in all of the species.
The details of variation in these and other characters and our
justifications for the synonymies are given in the accounts of the
species. All of the trivial names that apply to species herein
recognized as members of the genus _Osteocephalus_ are listed in
table 1.
=Osteocephalus= Steindachner, 1862
_Osteocephalus_ Steindachner, 1862:77 [Type
species.--_Osteocephalus taurinus_ Steindachner, 1862, by
original designation]. Not _Osteocephalus_ Fitzinger,
1843:50 (_nomen nudum_).
_Diagnostic Definition._--1) Skull broader than long; 2) dermal roofing
bones of skull well ossified, exostosed, and/or co-ossified in some
species; 3) prenasal and internasal bones absent; 4) parasphenoid alae
posterolaterally oriented; 5) dentigerous processes of prevomers angular
(/--\); 6) vocal sacs paired, posterior, and when inflated protruding
posteroventral or posterolateral to angles of jaws; 7) submentalis
muscle moderate in size and araphic; 8) intermandibularis muscle
undifferentiated and bearing an elongate median aponeurosis; 9)
parotoid glands absent or poorly developed, skin not producing viscous
secretion characteristic of _Phrynohyas_; 10) skin on dorsum tuberculate
in males, smooth in females; 11) tympanum large, 60 percent or more of
diameter of eye; 12) fingers about one-third, toes more than
three-fourths webbed; 13) discs large, round; 14) nuptial excrescences
present in breeding males; 15) inner metatarsal tubercle not modified
for digging; 16) outer metatarsal tubercle absent; 17) tarsal fold weak
or absent; 18) pupil horizontal; 19) palpebrum clear; 20) known tadpoles
having two upper and five lower rows of teeth.
TABLE 1.--Alphabetical Synonymy of the Species of _Osteocephalus_.
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