A Taxonomic Revision of the Leptodactylid Frog Genus Syrrhophus CopeLynch, John D.
Science
A Taxonomic Revision of the Leptodactylid Frog Genus Syrrhophus Cope
Lynch, John D.
Frogs; Syrrhophus
_Phyllobates cystignathoides_ Cope, 1877:89-90 [Syntypes.--USNM
32402-32409, from Potrero, near Cordoba, Veracruz, Mexico,
collected by Francis Sumichrast]. Boulenger, 1882:196.
_Syrrhophus cystignathoides_: Cope, 1879:268. Kellogg, 1932:
126-27. Taylor and Smith, 1945: 582-83. Smith and Taylor, 1948:50.
Martin, 1958:49.
_Syrrhaphus cystignathoides_: Guenther, 1900:218.
_Syrraphus cystignathoides_: Diaz de Leon, 1904:10.
_Syrrhopus cystignathoides_: Barbour and Loveridge, 1946:170.
[Illustration: FIG. 5: Distribution of _Syrrhophus cystignathoides
campi_ (solid symbols) and the nominate subspecies (open symbols).]
_Diagnosis._--Venter areolate; interorbital light and dark bars present;
ground color yellow to brownish-yellow in life (Fig. 4b).
_Remarks._--Firschein (1954) briefly considered the status of Peters'
(1871) _Phyllobates verruculatus_ and noted that if it was a
_Syrrhophus_ it would probably be referrable to _S. cystignathoides_.
Peters' (1871) original description corresponds well with
_S. cystignathoides_, and the type-locality ("Huanusco" = Huatusco) is
within the range of that species. Firschein (1954) expressed doubt that
_verruculatus_ was a _Syrrhophus_, because Peters placed it in another
genus. However, Peters described _verruculatus_ a decade before Cope
diagnosed the genus Syrrhophus. Most frogs now called _Syrrhophus_, plus
a number of lower Central American frogs now placed in a variety of
genera were placed in _Phyllobates_ by Boulenger, Cope, and Peters.
The types of _Phyllobates verruculatus_ were destroyed during World War
II (Guenther Peters, _in litt._); the specimens subsequently assigned to
the taxon by Kellogg (1932) are _Syrrhophus cystignathoides_. Because
the type specimens are lost and because the name antedates the more
established name, _cystignathoides_, I favor retaining _Phyllobates
verruculatus_ Peters as a _nomen dubium_.
Smith and Taylor (1948) reported _S. verruculatus_ from Tianguistengo,
Hidalgo, Mexico. These specimens are examples of _verrucipes_. Smith
(1947) reported a specimen of _verruculatus_ from San Lorenzo, Veracruz.
Firschein (1954) referred it to _cystignathoides_, and Duellman (1960)
concluded that both authors were in error and that the specimen (USNM
123530) was a _leprus_.
_Etymology._--The trivial name is the diminutive of _Cystignathus_, a
once-used generic name for several leptodactylid frogs.
_Distribution._--Low and moderate elevations in the foothills along the
Sierra Madre Oriental from eastern San Luis Potosi to Central Veracruz,
Mexico (Fig. 5).
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