Practically the whole of the path of this ameba consists of right and
left-hand curves which are nearly uniform in length, each wave being
about eight to ten times the length of the ameba. Since the drawings
were made at intervals of a minute, the waves are therefore from eight
to ten minutes long in time, measuring from crest to crest. Some of the
waves are flatter than others, for example wave No. 4, but otherwise it
is like the others. Wave 7 is a double wave due to a change of
direction. Instead of turning to the right at 9:23 the ameba changed its
direction and turned to the left. The smoothness with which this turn
was made indicates that it originated in the mechanism producing the
sinuous course itself, or that it proceeded from a very slight stimulus
external to it. At 9:49 the direction of movement was changed again, but
just enough to disturb the formation of a smooth wave. The general
direction of locomotion was not changed. It may be assumed that this
change was produced by a stimulus external to the wave-producing
mechanism. The irregularity and shortness of wave 13 was probably due to
the same stimulus that disturbed wave 11. Shortly after 9:58 the ameba
came within sensing range of a mass of debris which it pushed away and
followed, thus causing a change in the direction of movement. Although
waves begin to appear again after this, some of them very smooth, they
are not typical for they are too short, ranging from a little over three
lengths (wave 16) to a little over six lengths (wave 23). It is likely
that the disturbance caused by the mass of debris at 9:58 together with
the onset of the division crisis produced the succession of atypical
waves. An external disturbance that is sufficiently strong to change the
direction of locomotion usually persists for the duration of at least
one wave length thereafter. It will be noted that
[Illustration: Figure 33. Illustrating the path of an _Amoeba bigemma_
under controlled conditions, in ordinary diffuse light from a north
window. For convenience of reference the waves in the path are numbered
from 1 to 23. The figures were drawn with the aid of a camera lucida at
intervals of one minute. The path was recorded from 8:58 to 10:26, when
fission occurred. After fission, the path of one of the daughter cells
was followed for a short time, but the ingestion of a mass of debris
destroyed the regularity of the path. The vertical lines at the point
where fission occurred indicate that the figures _above_ the lines were
moved up the length of the lines. The first figure beyond wave 13 was
influenced in its movements by a mass of debris lying to its left.
Average length of the ameba, 150 microns.]
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