in _frenata_ in _erminea_
[M] ordinarily less than 46 [M] ordinarily more than 46
[F] less than 47 [F] more than 48
In the fossils the percentage for the larger skulls is 46; for the
smaller skulls it is 48.
It may be that the ten fossil skulls are six female _frenata_ and four
male _erminea_ but I think not. In the first place a skull of different
shape, seemingly of the _frenata_ stock, is known from the deposit and
it is almost certain that two subspecies of the same species would not
occur at the same place at the same time. It is possible, of course,
that parts of the deposits were laid down at times so far apart that a
shift in geographic range of two subspecies had occurred. This one
skull, seemingly of the _frenata_ stock, is the type of _Putorius
gracilis_ Brown (see p. 404) and was regarded as the only known
specimen of _gracilis_. Regardless of the specific identity of this one
specimen named _gracilis_, the chances of obtaining otherwise from a
deposit, like that in Conard Fissure, six females of frenata and four
males of _erminea_ without a male _frenata_ or a female of _erminea_
coming to light are so slight as strongly to incline me to the view
that the six larger specimens are males of the same species to which
the 4 smaller specimens belong. By either this interpretation, or the
one initially considered (of female _frenata_ and male _erminea_), the
animals from the fissure are at least subspecifically distinct from any
American Recent weasel. Furthermore, by this latter interpretation each
sex of this weasel, _angustidens_, is intermediate between the
_frenata_ and _erminea_ stocks in the feature of postglenoidal length
which feature, at any place where the two Recent species occur
together, serves to distinguish one from the other. In the northernmost
subspecies of _erminea_ (_arctica_ for example) the postglenoidal
length in some males is no longer than in males of _frenata_.
Considering general size, _angustidens_ agrees better with _erminea_
than with frenata and this circumstance has influenced me to place
_angustidens_ as a subspecies of _erminea_.
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