Today, _erminea_ is not known to occur nearer Conard Fissure than
northern Iowa, more than 400 miles to the northward. In comparison with
the race there, _bangsi_, males of _angustidens_ are of approximately
the same size but in the shorter distance between the glenoid fossa and
anterior margin of the tympanic bulla, and also in the lesser
postglenoidal length of the skull, _angustidens_ resembles the
northernmost American subspecies of _erminea_. Females of _angustidens_
differ more from any living weasel than the males do. The females are
much larger than those of _bangsi_, and among living American races of
_erminea_ most closely resemble intergrades between _arctica_ and
_richardsonii_ which intergrades are found approximately 1700 miles to
the north of Conard Fissure. In females, the preorbital part of the
skull in _M. e. arctica_ is broader and in _M. e. richardsonii_
narrower than in _angustidens_. If it seems strange that females of
_angustidens_ resemble one subspecies whereas males, in size, resemble
another subspecies almost a thousand miles distant, it should be
remembered that the degree of sexual dimorphism varies much from one
subspecies to another in the Recent animals. An example is furnished by
_Mustela erminea fallenda_ and _Mustela erminea invicta_.
The assemblage of mammals from Conard Fissure includes several species
of boreal predilections which, like _Mustela erminea_, now occur only
much farther north than Arkansas. At one time the edge of the sheet of
ice was only about 200 miles north of Arkansas. It may be significant
that the cranial characters of the female ermine from the Fissure, and
qualitative cranial characters of males from there, are most nearly
approximated among Recent weasels by those which live along the
southern edge of the frozen tundra.
In view of what has been said, the possibility should be considered
that the distinctive cranial features of _angustidens_ may be the
result of evolutionary change in time as well as of geographic
variation resulting from horizontal placement.
=MUSTELA RIXOSA= (Bangs)
Least Weasel
(Synonymy under subspecies)
_Type._--_Putorius rixosus_ Bangs, Proc. Biol. Soc. Washington,
10:21, February 25, 1896.
_Range._--From Norway and Switzerland eastward through Siberia and
all the way across North America, but unknown from Iceland,
Greenland and the Arctic islands west of Greenland; in North
America, from the Arctic Life-zone south to Central British
Columbia, Montana and into parts of the Upper Austral Life-zone as
in the eastern half of the continent.
Public-domain text, read in full here on John Shaqi.
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