But now, in the second place, not only is this assumption wholly
gratuitous, but there are many considerations which render it in the
highest degree improbable, while there are not wanting facts which
appear to demonstrate that it is false. For, unquestionably, most of
the considerations which have already been advanced in the preceding
chapter against the assumption of an absolute stability of germ-plasm,
are here equally available against the assumption of an imperceptibly
small amount of instability[63]. Similarly, all the facts there given
with regard to the a-sexual origin of species—and even genera—of
parthenogenetic organisms, bud-variation[64], &c., amply demonstrate
that congenital variations due to the instability of germ-plasm alone,
or apart from amphimixis, are sometimes enormous. Hence, we cannot
accept the gratuitous suggestion that in all other cases they are too
insignificant to count for anything till they have been augmented by
amphimixis, even although we may be prepared to agree that amphimixis
is probably one important factor in the production of congenital
variations. What degree of importance it presents in this connexion,
however, we have not at present any means of determining; all we can
conclude with certainty is, that in some cases it is demonstrably very
much less than Weismann supposes, while it is extremely improbable
that it is ever in any case the sole and necessary antecedent to the
operation of natural selection.
This extreme improbability is shown, not only by what I have already
said in the previous chapter, and need not here repeat; but likewise
by the “several considerations” which Darwin has adduced with regard
to this very point, and which, as he says, “alone render it probable
that variability of every kind is directly or indirectly caused by
changed conditions of life,” with the consequence that “those authors
who attribute all variability to the mere act of sexual union are in
error.” I have already quoted these words further back in the present
chapter, in order to show that by now attributing the _origin_ of all
congenital variations to the direct action of external conditions,
Weismann has brought himself into line with Darwin so far as this
fundamental point of doctrine is concerned. But I here re-quote the
words in order to show that by further attributing the _development_ of
congenital variations “to the mere act of sexual union,” Weismann is
again falling out of line with Darwin. So to speak, he first performs a
right-about-face movement as regards his original position towards the
“stability of germ-plasm,” and immediately afterwards makes a half-turn
back again. Now, it is this half-turn to which I object as unwarranted
in logic and opposed to fact.
Public-domain text, read in full here on John Shaqi.
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