In a previous chapter (pp. 66-7) I presented to him the dilemma,
that germ-plasm must be either absolutely stable or else but highly
stable, and that in the former case his theory of amphimixis as the
sole cause of congenital variations would be valid, while in the
latter case the theory would collapse. But it did not then occur to
me that Weismann might seek a narrow seat between the horns of this
dilemma, by representing that germ-plasm is universally unstable up to
a certain very low degree of instability—viz., exactly that degree
which is required for starting a congenital variation by means of
external causes, without its being possible for the variation to become
perceptible unless afterwards increased by means of amphimixis. And now
that this extremely sophistical position has been adopted, I cannot
see any imaginable reason for adopting it other than a last endeavour
to save as much as possible of his former theory of evolution. There
can be nothing in the nature of things thus to limit, within the
narrowest possible range, the instability of a universally unstable
germ-plasm—distributed, as this most complex of known substances
is, throughout all species of plants and animals, and exposed to
inconceivably varied conditions of life in all quarters of the globe.
And these considerations are surely of themselves enough to dispose
of the assumption as absurd, without again rehearsing the facts of
congenital variation which definitely prove it to be false.
_Conclusion._
For reasons stated at the commencement of this chapter, I have
restricted its subject-matter almost exclusively to a consideration
of the more fundamental changes which Professor Weismann has wrought
in his general system of theories by the publication of his most
recent works. In other words, I have purposely avoided considering
those immensely elaborate additions to his theory of heredity which
constitute by far the largest portion of his essays on _Amphimixis_ and
_The Germ-plasm_, and which have for their object an ideal construction
of “the architecture of germ-plasm.”
The fundamental changes to which allusion has just been made are as
follows.
Professor Weismann has to a large extent abandoned his theory of polar
bodies, and in my opinion would have done well had he taken a further
step and surrendered the theory _in toto_.
Public-domain text, read in full here on John Shaqi.
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