But let us neglect the probability of my hypothesis, and consider
merely its logical accuracy. Professor Vines says:—“The fate of the
germ-plasm of the fertilized ovum is, according to Professor Weismann,
to be converted in part into the somato-plasm (!) of the embryo, and
in part to be stored up in the germ-cells of the embryo. This being
so, how are we to conceive that the germ-plasm of the ovum can impress
upon the somato-plasm (!) of the developing embryo the hereditary
character of which it (the germ-plasm) is the bearer? This function
cannot be discharged by that portion of the germ-plasm of the ovum
which has become converted into the somato-plasm (!) of the embryo,
_for the simple reason that it has ceased to be germ-plasm_, and must
therefore have lost the properties characteristic of that substance.
Neither can it be discharged by that portion of the germ-plasm of the
ovum which is aggregated in the germ-cells of the embryo, for under
these circumstances it is withdrawn from all direct relation with the
developing somatic-cells. The question remains without an answer.” I
believe myself to have answered this above. I do not recognize the
somato-plasm of Professor Vines; my germ-plasm, or idio-plasm of the
first ontogenetic grade, is not modified into the somato-plasm of
Professor Vines, but into idio-plasm of the second, third, fourth,
hundredth, &c. grade, and every one impresses its character on the
cell containing it.
It may be dullness, but I confess that this does not appear to me an
“answer” to Professor Vines’ criticism. Even though “idio-plasm of
the first ontogenetic grade” has to become “idio-plasm of the second,
third, fourth, hundredth, &c. grade,” before in each of the grades
concerned it can give origin to the somatic-cells which are distinctive
of that grade, I cannot see that it makes any difference (in relation
to Vines’ criticism) whether we speak of those cells as containing
“somato-plasm,” or as containing “somatic idio-plasm” of such and such
a grade, _plus_ “cytoplasm.” For whether we thus follow Weismann’s
earlier terminology or his later, we are so far speaking about exactly
the same thing, namely, the transformation of “germ-plasm” into all
the constituent cells of the “soma.” The difficulty is, in Vines’
words above cited, “to conceive that the germ-plasm of the ovum can
impress upon the somato-plasm of the developing embryo the hereditary
characters of which it (the germ-plasm) is the bearer”; and Weismann
says that this difficulty, which he acknowledges, can now be answered
by substituting for his original statement that “germ-plasm” becomes
changed into “somato-plasm,” the statement that it is “idio-plasm”
_derived_ from “germ-plasm” which thus “impresses its character on the
cell containing it.” But, “as a matter of logical accuracy,” there
is surely here a distinction without a difference. For what is the
difference between saying that germ-plasm “impresses” its character
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