Hence, the nuclear contents (nucleo-plasm) of every cell, whether
germinal or somatic, present two substances, which we may, in the
absence of any better terms supplied by Weismann himself, respectively
designate “idio-plasm-A” and “idio-plasm-B.” Idio-plasm-A is the
substance which has to do only with the formation and control of the
individual cell in which it resides, like a mollusc in its shell.
Idio-plasm-B is the substance out of which future cells are to be
formed and controlled, when in due course either of ontogeny or
phylogeny this idio-plasm-B becomes converted into idio-plasm-A,—i.e.,
into each subsequently developing tissue or organism, as the case may
be. I say ontogeny or phylogeny, and tissue or organism, because, where
a _germ-cell_ is concerned, idio-plasm-B is capable of reproducing
entire organisms of its own and of subsequent generations; whereas,
in the case of all _somatic_-cells, idio-plasm-B is capable only
of reproducing, stage by stage, some greater or less number of the
cells which are to construct the single organism of which they form a
part. Or, otherwise expressed, in the particular case of a germ-cell
idio-plasm-B is germ-plasm, and therefore is alone capable of producing
an entire organism of somatic-cells, while it is likewise alone capable
of reproducing successive organisms; for it alone contains the carriers
of heredity[7].
Thus, idio-plasm-B of an unsegmented germ-nucleus is germ-plasm. But
as soon as the germ-nucleus has undergone its first nuclear division,
its nucleo-plasm is no longer germ-plasm, inasmuch as each of the
half-portions is now no longer capable of reproducing an entire
organism—unless it be in the case of identical twins. Similarly in
the second nuclear division, each of the four resulting idio-plasms-B
is still further removed from the pristine character of germ-plasm;
and so on through all successive stages of segmentation. Hence
these successive nuclear divisions must indicate a partitioning and
re-partitioning of the original idio-plasm-B (germ-plasm) into the
idio-plasms-B severally distinctive of all the various cells of the
soma.
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