afterwards called germ-plasm, in that it always occupied a separate
“sphere” of its own, where its continuity has been uninterrupted “since
the first origin of life.” But Galton was not seduced by the temptation
to construct an ideally logical system; and he had what I regard as the
sound judgement to abstain from carrying his theory of stirp into any
such transcendental “sphere” as that which is occupied by Weismann’s
theory of germ-plasm, in relation to the general doctrine of descent.
* * * * *
There is, then, a vast distinction between any theory of heredity
which postulates the material of heredity as highly stable and largely
continuous, and Weismann’s theory, which postulates this material as
absolutely stable and perpetually continuous. But we must next take
notice that Weismann himself has not kept this distinction in view
with the constancy which we should have expected from so forcible a
thinker. On the contrary, although in the construction of his theory
of evolution he never fails to press the postulates of _absolute_
stability and _perpetual_ continuity to their logical conclusions in
the various doctrines above enumerated (pp. 57-58), when engaged on
his more special theory of heredity he every now and then appears to
lose sight of the distinction. Indeed, he occasionally makes such large
concessions with regard to both these postulates, that, were they to be
entertained, the occupation of his critics would be gone: his theory
of heredity would become converted into Galton’s, while his theory of
evolution would vanish altogether. It is therefore necessary to quote
some of these concessions, if only to justify ourselves in subsequently
ignoring them. I will give one instance of each; but it is necessary to
preface the illustrations with a few words to mark emphatically three
very distinct cases of congenital variation—leaving aside for the
present the question whether or not they all occur in fact, as they are
held to do by one or other of the theories of heredity.
1. The case where impoverished nutrition of the body has the effect
of simply _starving_ its germinal material. This is not a case
where either the continuity or the stability of such material is
affected. Its full efficiency as “formative material” may indeed be
thus deteriorated to any extent, so that the progeny may be to any
extent puny or malformed; but this will not necessarily cause any
such re-shuffling of its “molecules” as will thereafter result in
a permanent phylogenetic change. At most it will affect only the
immediate offspring of poorly nourished parents; and natural selection
will always be ready to eliminate such inefficient individuals. This
case I will always hereafter call the case of _nutritive_ congenital
changes.
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