It will be perceived that Weismann himself here very clearly draws
all the distinctions between cases 1, 2, and 3, as above explained.
Therefore it becomes the more remarkable that he should not have
perceived how radically inconsistent it is in him thus to entertain
as “possible” congenital variations belonging to the case 2. For, as
we have now so fully seen, the theory of germ-plasm (as distinguished
from that of stirp) cannot entertain the possibility of an hereditary
and specialized change of _any_ kind as thus produced by external
conditions of life: should such a possibility be entertained, there
must obviously be an end to the _absolute stability_ of germ-plasm,
and a consequent collapse of Weismann’s theory of evolution. Either
germ-plasm is absolutely stable, or else it is but highly stable. If
it is absolutely stable, individual variations of an hereditary kind
can occur only as results of sexual admixtures of germ-plasm, and
Weismann’s theory of evolution is established. But if germ-plasm is
not absolutely stable (no matter in how high a degree it may be so)
hereditary individual variations may be produced by other causes, and
Weismann’s theory of evolution collapses. Therefore, if we are to
examine his theory of _evolution_, we can do so only by ignoring such a
passage as the one just quoted, which surrenders the postulate of the
_absolute stability_ of germ-plasm.
Again, if we are to examine Weismann’s theory of _heredity_, we must
similarly ignore such a passage as the following, where he represents
that he is similarly prepared to surrender his still more fundamental
postulate of the _perpetual continuity_ of germ-plasm.
After remarking that some of his own experiments on the climatic
varieties of certain butterflies raise such difficulties against his
whole theory of heredity that even now “he cannot explain the facts
otherwise than by supposing the passive acquisition of characters
produced by the direct influence of climate,” he goes on to remark more
generally—“We cannot exclude the possibility of such a transmission
occasionally occurring, for, even if the greater part of the effects
must be attributed to natural selection, there might be a smaller part
in certain cases which depends on this exceptional factor[19]”—i.e.,
the Lamarckian factor!
Public-domain text, read in full here on John Shaqi.
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