5. That a large proportional number of the gemmules in each packet,
however, fail to develop, and are then transmitted in a dormant
state to future generations, in any of which they may be developed
subsequently—thus giving rise to the phenomena of reversion or atavism.
6. That in all cases the development of gemmules into the form of their
parent cells depends on their suitable union with other partially
developed gemmules, which precede them in the regular course of growth.
7. That gemmules are thrown off by all physiological cells, not only
during the adult state of the organism, but during all stages of
its development. Or, in other words, that the production of these
cell-seeds depends upon the adult condition of parent cells: not upon
that of the multicellular organism as a whole.
At first sight it may well appear that we have here a very formidable
array of assumptions. But Darwin ably argues in favour of each of them
by pointing to well-known analogies, drawn from the vital processes
of living cells both in the protozoa and metazoa. For example, it
is already a well-recognized doctrine of physiology that each cell
of a metazoon, or multicellular organism, though to a large extent
dependent on others, is likewise to a certain extent independent
or autonomous, and has the power of multiplying by self-division.
Therefore, as it is certain that the sexual elements (and also buds
of all descriptions) include formative material of some kind, the
first assumption—or that which supposes such formative matter to be
participate—is certainly not a gratuitous assumption.
Again, the second assumption—viz., that this particulate and formative
material is dispersed throughout all the tissues of the organism—is
sustained by the fact that, both in certain plants and in certain
invertebrated animals, a severed portion of the organism will develop
into an entire organism similar to that from which it was derived, as,
for example, is the case with a leaf of Begonia, and with portions cut
from certain invertebrated animals, such as sea-anemones, jelly-fish,
&c. This well-known fact in itself seems enough to prove that the
formative material in question must certainly admit, at all events in
many cases, of being distributed throughout all the tissues of living
organisms.
The third assumption—or that which supposes the formative material
to be especially aggregated in the sexual elements—is not so much an
assumption as a statement of obvious fact; while the fourth, fifth,
sixth, and seventh assumptions all follow deductively from their
predecessors. In other words, if the first and second assumptions be
granted, and if the theory is to comprise all the facts of heredity,
then the remaining five assumptions are bound to follow.
Public-domain text, read in full here on John Shaqi.
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