Similarly as regards his postulate of the stability of germ-plasm
as _absolute_. It is enough for all the requirements of his theory
of heredity, that the substance in question should present the high
degree of stability which the facts of atavism, persistence of
vestigial organs, &c., prove it to possess. But for his further
theory of evolution it is necessary to make this further postulate of
the stability of germ-plasm _as undisturbed since the first origin of
sexual propagation_: otherwise there would be no logical foundation for
any of the distinctive doctrines which go to constitute that theory.
Thus much understood, we proceeded to examine the theory of germ-plasm
in each of its departments separately—i.e., first as a theory of
heredity, and next as a theory of evolution. And we begun by comparing
it as a theory of heredity with the preceding theories of Darwin and
Galton. In the result we found that germ-plasm resembles gemmules in
all the following respects. It is particulate; constitutes the material
basis of heredity; is mainly lodged in highly specialized cells; is
nevertheless also distributed throughout the general cellular tissues,
where it is concerned in all processes of regeneration, repair, and
a-sexual reproduction; presents an enormously complex structure,
in that every constituent part of a potentially future organism is
represented in a fertilized ovum by corresponding particles; is
everywhere capable of virtually unlimited multiplication, without ever
losing its hereditary endowments; is often capable of carrying these
endowments in a dormant state through a long series of generations,
until at last they reappear again in what we recognize as reversions.
Such being the points of resemblance, the only points of difference
may be summed up in the two words—continuity, and stability. For,
as regards continuity, while Darwin’s theory supposes the substance
of heredity to be more or less formed anew in each generation by
the body-tissues of that generation, Weismann’s theory regards
this substance as owing nothing to the body-tissues, further than
lodgement and nutrition. Therefore, while the theory of gemmules can
freely entertain the doctrines of Lamarck, the theory of germ-plasm
excludes them as physiologically impossible, in all cases where
sexual reproduction is concerned. Again, as regards stability, while
Darwin’s theory simply accepts the fact of such a degree of stability
appertaining to the substance of heredity as the phenomena of atavism,
&c. prove, Weismann’s theory postulates the stability of this substance
as absolute. But, as we have now so often seen, he does so in order to
provide a hypothetical basis for his further theory of evolution. In as
far as his theory of heredity is concerned, there is no reason why it
should differ from Darwin’s in this respect.
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