Again, comparing Weismann’s theory of heredity with that of Galton, we
found that germ-plasm resembles stirp in all the points wherein we have
just seen that it resembles germ-plasm. Or, otherwise stated, all three
theories are thus far coincident. But germ-plasm resembles stirp much
more closely than it does gemmules, seeing that the theory of stirp is
founded on the postulate of “continuity” in exactly the same manner
as is the theory of germ-plasm. In point of fact, the only difference
between these two theories consists in the two further postulates
presented by the latter—viz., that the “continuity” in question has
been unbroken since the origin of life, while the “stability” in
question has been uninterrupted since the origin of sexual propagation.
But seeing that both these additional postulates have reference to
Weismann’s theory of evolution, we may say that his theory of heredity
is, as regards all essential points, indistinguishable from that of
Galton.
The truly scientific attitude of mind with regard to the problem of
heredity is to say, as Galton says, “that we might almost reserve our
belief that the structural [i.e., somatic] cells can react on the
sexual elements at all, and we may be confident that at most they do
so in a very faint degree; in other words, that acquired modifications
are barely, if at all, _inherited_, in the correct sense of that word.”
But for Weismann’s further theory of evolution, it is necessary to
postulate the two additional doctrines in question; and it makes a
literally immeasurable difference to the theory of evolution whether
or not we entertain these two additional postulates. For no matter how
faintly or how fitfully the substance of heredity may be modified by
somatic tissues, by external conditions of life, or even by so-called
spontaneous changes on the part of this substance itself, numberless
causes of congenital variation are thus admitted, while even the
Lamarckian principles are hypothetically allowed some degree of play.
And although this is a lower degree than Darwin supposed, their
influence in determining the course of organic evolution may still have
been enormous; seeing that their action in any degree must always have
been _directive_ on the one hand, and _cumulative_ on the other.
Public-domain text, read in full here on John Shaqi.
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