The next most important point of difference between the theories of
stirp and germ-plasm has reference to the mechanism of ontogeny.
According to Galton, this is simply a struggle between all the
carriers of heredity composing the stirp of a fertilized ovum. It
is not, however, a struggle for existence, but what may be called a
struggle for development. In the fertilized ovum all the carriers
of heredity are, to begin with, in a “latent” condition; but of
this enormous multitude of “germs” or “gemmules,” only a very small
proportional number are destined to become “patent”—i. e., developed
into the tissue-cells composing the new organism. The vast majority
of the gemmules, or those which fail to be thus developed, go to
constitute the stirp of the new organism when this has been formed by
the development of the comparatively few successful gemmules. Thus much
understood, the following quotation will be fully intelligible.
My argument is this: Of the two groups of germs, the one consisting
of those that succeed in becoming developed and in forming the bodily
structure, and the other consisting of those that remain continually
latent, the latent vastly preponderates in number. We should expect
the latent germs to exercise a corresponding predominance in matters
of heredity, unless it can be shown that, on the whole, the germ that
is developed into a cell becomes thereby more fertile than if it had
remained latent. But the evidence points the other way. It appears
both that the period of fertility is shorter, and the fecundity even
during that period is less in the germ that becomes developed into a
cell, than they are in the germ that remains latent. Much less then
would the entire bodily structure, which consists of a relatively
small number of these comparatively sterile units, successfully
compete in matters of heredity with the total effect of the much more
numerous and more prolific units which are in a latent form[46].
Thus, Galton’s theory of the mechanism of geny is a theory of
struggle; and this constitutes a point of difference on which Weismann
lays much stress in his latest work. For, as we know, Weismann regards
the mechanism of ontogeny as characterized by a peaceful succession
of “stages,” which are “pre-determined from the germ onwards”; and
in his latest work this idea of orderly sequence has been further
elaborated in his doctrine of “determinants.” In short, to adopt
their own metaphors, while Galton tells us that the mechanism of
ontogeny is like that of a political election, where rival candidates
compete to “represent” the nation (stirp) in Parliament (individual
organism); Weismann likens it to the mechanism of a well-drilled army,
where ultimate carriers of heredity (privates) are banded together
in companies, regiments, battalions, &c., under the command of
corresponding officers (determinants).
Public-domain text, read in full here on John Shaqi.
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