Lastly, there is yet one further point of difference between stirp and
germ-plasm, which is thus stated by Weismann:—
Galton’s idea is only conceivable on the presupposition of the
occurrence of sexual reproduction, while the theory of the continuity
of the germ-plasm is entirely independent of any assumption as to
whether each primary constituent is present in the germ _singly_
or in numbers. According to my idea, the active and the reserve
germ-plasm contain precisely similar primary constituents, gemmules,
or determinants; and on this the resemblance of a child to its
parent depends. The theory of the continuity of the germ-plasm, as I
understand it, is not based on the fact that each “gemmule” necessary
for the construction of the soma is present many times only, so that
a residue remains from which the germ-cells of the next generation
may be formed: it is founded on the view of the existence of a
special adaptation, which is inevitable in the case of multicellular
organisms, and which consists in the germ-plasm of the fertilized
egg-cell becoming doubled primarily, one of the resulting portions
being reserved for the formation of germ-cells[47].
These being the main points of difference between the theories of stirp
and of germ-plasm to which Professor Weismann has alluded, I will now
proceed to consider them separately, in reverse order to that in which
they have been here stated.
The point of difference last mentioned need not detain us long, because
it seems to me one of very little importance. “Whether each primary
constituent is present in the germ singly or in numbers” cannot greatly
signify, so long as both theories agree that, sooner or later, they
must be present plurally. Galton supposes them to be thus present
from the first (i. e. in the unfertilized ovum), while Weismann
supposes them to be so only as a result of their self-multiplication
at a somewhat later stage (i. e. in the segmenting ovum, and onwards
throughout the procreative life of the individual). Doubtless Weismann
does not suppose that they ever become so numerous as Galton imagines;
but the whole question is so highly speculative that I do not see how
any useful purpose can be served by debating it. Nor do I see why
Weismann should conclude that “Galton’s idea is only conceivable on the
presupposition of the occurrence of sexual reproduction.” It is true
that Galton has discussed exclusively the case of sexual reproduction;
but I cannot perceive that any of his ideas are inapplicable to
a-sexual.
Public-domain text, read in full here on John Shaqi.
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