An introduction to the study of fishesGünther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Science
An introduction to the study of fishes
Günther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Fishes; Ichthyology
The structure of the intestinal tract of _Teleosteous_ fishes is
subject to so numerous modifications that we should go beyond the
limits of the present work if we would attempt to enter into details.
Great differences in this respect may be found even in groups of the
same natural families. Frequently the intestinal tract remains of
nearly the same width throughout its course, and only the entrance
of the various ducts serves as a guide for the distinction of its
divisions. An intestine of such uniform width may be straight and
short, as in _Scombresocidæ_, _Symbranchidæ_, or it may be more or less
convoluted and long, as in many _Cyprinidæ_, _Doradina_, etc. On the
whole, carnivorous fishes have a much shorter and simpler intestinal
tract than herbivorous.
In the majority of Teleosteans, however, œsophagus, stomach, duodenum,
small intestine and rectum, can be more or less distinctly, even
externally distinguished.
There are two predominant forms of the stomach, intermediate forms
being, however, numerous. In the first, the _siphonal_, it
presents the form of a bent tube or canal, one-half of the horse-shoe
being the cardiac, the other the pyloric portion. In the second, the
_cæcal_, the cardiac division is prolonged into a long descending
blind sac, the cardiac and pyloric openings of the stomach lying close
together (_Clupea_, _Scomber_, _Thynnus_, etc.)
[Illustration: Fig. 56.--Siphonal Stomach and Pyloric Appendages
of a Female Salmon, 3⅓ feet long. _a a a_, Pyloric appendages;
_ch_, ductus choledochus; _oe_, œsophagus; _st_, lower end
of stomach; _p_, pyloric region; _i_, ascending; and _í’_,
descending portion of intestine.]
The duodenum receives always the hepatic and pancreatic secretions,
and, besides, those of the appendices pyloricæ, which, in varying
numbers (from 1 to 200), are of very common occurrence in Teleosteans
(Fig. 56). They vary also in length and width, and whilst the
narrowest serve only as secretory organs, the widest are frequently
found filled with the same contents as the intestine. When few in
number, each opens by a separate duct into the duodenum; when their
number is greater two or more coalesce into a common duct; in the
latter case the appendages cease to be free, and are connected with one
another by a more or less firm tissue.
Cœcal appendages at the end of the intestinal tract are of exceedingly
rare occurrence in fishes (_Box_). There is no _cloaca_ in Teleosteans.
In the majority of Teleosteous fishes the _vent_ is situated on the
boundary between trunk and tail, behind the ventral fins. In a few it
lies farther backwards, not far from the caudal fin; more frequently
it is advanced forwards, under the middle of the abdomen or to the
scapular arch. In two fishes, _Aphredoderus_ and _Amblyopsis_, it lies
before the pectoral fins.
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