An introduction to the study of fishesGünther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Science
An introduction to the study of fishes
Günther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Fishes; Ichthyology
A peritoneum envelops all the divisions of the intestinal tract within
the abdominal cavity. A broad, well-developed _omentum_ has hitherto
been found in _Gobiesox cephalus_ only.
_Liver_.--The existence of a liver in _Branchiostoma_ as a long
diverticulum of the intestine has been mentioned above. In the
Myxinoids the liver is divided into two glandular bodies, an anterior
rounded smaller one, and a posterior larger one of an elongate shape.
The gall-bladder lies between both, and receives a cystic duct from
each of them. In the other fishes the proportionally large liver is a
single large gland, from which only now and then small portions are
found to be detached. It is either simple, or with a right and left
lobe, or with a third lobe in the middle; each lobe may have incisions
or subdivisions, which, however, are very inconstant. The liver of
fishes is distinguished by the great quantity of fluid fat (oil) which
it contains. The gall-bladder is but rarely absent, and attached to
the right lobe, or towards the centre; however, in some fishes it
is detached from the liver and connected with it by the cystic duct
only. The bile may be conveyed by one or more hepatic ducts into a
common duct which is continued towards the gall-bladder as _ductus
cysticus_, and towards the duodenum as _ductus choledochus_; or some
of the hepatic ducts enter directly the gall-bladder, or directly
the duodenum, without communicating with the common duct. Individual
variations in this respect are of common occurrence.
A _pancreas_ has been found hitherto in all Chondropterygians,
Acipenser, and many Teleosteans. In the first it is a glandular mass
of considerable size behind the stomach, close to the spleen; its duct
leads into the duodenum. In the Sturgeons the pancreas is attached to
the duodenum, and opens close to the ductus choledochus. In _Silurus
glanis_ it is very large, and the ductus choledochus passes through
its substance; it is smaller in _Belone_ and _Pleuronectes_, and
situated in the mesentery; its duct accompanies the terminal portion
of the ductus choledochus. In the Salmon, which possesses a large
lobed pancreas, the duct is so intimately connected with the ductus
choledochus that both appear externally as a single duct only.
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