An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
_Halitherium._[121]—The Miocene and early Pliocene seas of Europe
abounded in Sirenians, to which the generic name of _Halitherium_ was
given by Kaup, but which have also received other names. They had large
tusk-like incisors in the upper jaw, as in the existing Dugongs, though
not so greatly developed. Their molar teeth were ⁵⁄₅ or ⁶⁄₆, anteriorly
simple and single-rooted, posteriorly those above with three and those
below with two roots, and with enamelled and tuberculated or ridged
crowns, in all which respects they more resemble those of the Manatee
than of the Dugong. The anterior molars were deciduous; and there is
evidence of the presence of milk-teeth. Germs of inferior incisors were
also present. Some species at least had nasal bones, short, broad, but
normal in position, whereas in all the existing genera these bones
are quite rudimentary. Another and still more important evidence of
conformity to the general mammalian type is the better development of
the pelvic bone, and the presence of a small styliform femur articulated
to the acetabulum, although no traces of any other part of the limb
have been discovered. These ancient Sirenians, which may be regarded
as representing a distinct family—_Halitheriidæ_—were thus, in dental,
cranial, and other osteological characters, less specialised than are
either of the existing species, and if the intermediate links could be
discovered might well be looked upon as the ancestral forms from which
the latter have been derived, but at present the transitional conditions
have not been detected. So far as is yet known, when changes in the
physical conditions of the European seas rendered them unfitted to be the
habitation of Sirenians, the _Halitherium_ type still prevailed. If the
existing Dugongs and Manatees are descended from it, their evolution must
have taken place during the Pliocene and Pleistocene epochs, the one in
seas to the east, the other to the west of the African continent, which
has long formed a barrier to their intercommunication. _Halitherium_
remains have been found in many parts of Germany, especially near
Darmstadt, also in France, Italy, Belgium, Malta, etc. Until a few years
ago none were known from England, probably owing to the absence of beds
of an age corresponding to those in which they are found on the European
continent; but a skull and several teeth have been detected among the
rolled debris of which the Red Crag of Suffolk is partially composed. The
species are not yet satisfactorily characterised. Some of them appear
to have attained a larger size than the existing Manatee or Dugong. One
of these, from the Pliocene of Italy and France, having but ⁵⁄₅ molar
teeth, has been separated generically under the name of _Felsinotherium_
by Capellini, by whom it has been fully described; but the difference in
the number of the teeth does not afford sufficient grounds for separation
from _Halitherium_. _Miosiren_ of the Belgian Miocene, differs in that
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