An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
Teeth never functionally developed, but always disappearing before the
close of intra-uterine life. Palate provided with plates of baleen
or “whalebone.” Skull symmetrical. Nasal bones forming a roof to the
anterior nasal passages, which are directed upwards and forwards.
Maxilla produced in front of, but not over, the orbital process of the
frontal. Lachrymal bones small and distinct from the jugal. Tympanic
bone involuted (Fig. 76), and ankylosed with the periotic, which is
attached to the base of the cranium by two strong diverging processes.
Olfactory organ distinctly developed. Rami of mandible arched outwards,
their anterior ends meeting at an angle, and connected by fibrous tissue
without any true symphysis. All the ribs at their upper extremities
articulating only with the transverse processes of the vertebræ; their
capitular processes, when present, not articulating directly with
the bodies of the vertebræ. Sternum composed of a single piece, and
articulating only with a single pair of ribs. No ossified sternal ribs.
External openings of nostrils distinct from each other, longitudinal. A
short conical cæcum.
These animals have, when in the fœtal state, numerous minute calcified
teeth lying in the dental groove of both upper and lower jaws. They are
best developed about the middle of fœtal life, after which period they
are absorbed, and no trace of them remains at the time of birth.[129] The
baleen or whalebone does not make its appearance until after birth. It
consists of a series of flattened horny plates, between three and four
hundred in number, on each side of the palate, with a bare interval along
the middle line. These plates are placed transversely to the long axis of
the palate, with very short intervals between them. Each plate or blade
is somewhat triangular in form, with the base attached to the palate
and the apex hanging downwards. The outer edge of the blade is hard and
smooth; but the inner edge and apex fray out into long bristly fibres,
so that the roof of the Whale’s mouth looks as if covered with hair, as
described by Aristotle. At the inner edge of each principal blade are
two or three much smaller or subsidiary blades. The principal blades are
longest near the middle of the series, and gradually diminish towards the
front and back of the mouth. The horny plates grow from a dense fibrous
and highly vascular matrix, covering the palatal surface of the maxillæ,
and sending out lamellar processes, one of which penetrates the base of
each blade. Moreover, the free edge of these processes is covered with
very long vascular thread-like papillæ, one of which forms the central
axis of each of the hair-like epidermic fibres of which the blade is
mainly composed. A transverse section of fresh whalebone shows that it is
made up of numbers of these soft vascular papillæ, circular in outline,
each surrounded by concentrically arranged epidermic cells, and the whole
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