An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
In the most typically herbivorous forms of dentition, as seen in the
Horse and Kangaroo, the incisors are well developed, trenchant, and
adapted for cutting off the herbage on which the animals feed; the
canines are rudimentary or suppressed; the molars are large, with broad
crowns, which in the simplest forms have strong transverse ridges, but
may become variously complicated in the higher degrees of modification
which this type of tooth assumes.
Various forms of teeth of this type will be noticed among the Ungulates
and Rodents.
The natural groups of mammals, or those which in our present state of
knowledge we have reason to believe are truly related to each other, may
each contain examples of more than one of these modifications. Thus the
Primates have both omnivorous and insectivorous forms. The Carnivora show
piscivorous, carnivorous, insectivorous, and omnivorous modifications of
their common type of dentition. The Ungulata and the Rodentia have among
them the omnivorous and various modifications, both simple and complex,
of the herbivorous type. The Marsupialia exhibit examples of all forms,
except the purely piscivorous. Other orders, more restricted in number or
in habits, as the Proboscidea and Cetacea, naturally do not show so great
a variety in the dental structure of their members.
_Taxonomy._—In considering the taxonomic value to be assigned to the
modifications of teeth of mammals, two principles, often opposed to each
other, which have been at work in producing these modifications, must
be held in view:—(1) the type, or ancestral form, as we generally now
call it, characteristic of each group, which in most mammals is itself
derived from the still more generalised type described above; and (2)
variations which have taken place from this type, generally in accordance
with special functions which the teeth are called upon to fulfil in
particular cases. These variations are sometimes so great as completely
to mask the primitive type, and in this way the dentition of many animals
of widely different origin has come to present a remarkable superficial
resemblance, as in the case of the Wombat (a Marsupial), the Aye-Aye (a
Lemur), and the Rodents, or as in the case of the Thylacine and the Dog.
In all these examples indications may generally be found of the true
nature of the case by examining the earlier conditions of dentition;
for the characters of the milk-teeth or the presence of rudimentary
or deciduous members of the permanent set will generally indicate the
route by which the specialised dentition of the adult has been derived.
It is perhaps owing to the importance of the dental armature to the
well-being of the animal in procuring its sustenance, and preserving
its life from the attacks of enemies, that great changes appear to have
taken place so readily, and with such comparative rapidity, in the forms
of these organs—changes often accompanied with but little modification
in the general structure of the animal.
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