An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
In the Armadillos and their extinct allies alone is there an ossified
exoskeleton, or bony covering developed in the skin. In all other mammals
the skeleton is completely internal. It may be described as consisting
of an axial portion belonging to the head and trunk, and an appendicular
portion belonging to the limbs. There are also certain bones called
splanchnic, being developed within the substance of some of the viscera.
Such are the _os cordis_ and _os penis_ found in some mammals.
It is characteristic of all the larger bones of the mammalia that their
ossification takes its origin from several distinct centres. One near
the middle of the bone, and spreading throughout its greater portion,
constitutes the _diaphysis_, or “shaft,” in the case of the long
bones. Others near the extremities, or in projecting parts, form the
_epiphyses_, which remain distinct during growth, but ultimately coalesce
with the rest of the bone.
_Axial skeleton._—The axial skeleton consists of the skull, the vertebral
column (prolonged at the posterior extremity into the tail), the sternum,
and the ribs.
_Skull._—In the _skull_ of adult mammals, all the bones, except the
lower jaw, the auditory ossicles, and the bones of the hyoid arch, are
immovably articulated together, their edges being in close contact,
and often interlocking by means of fine denticulations projecting from
one bone and fitting into corresponding depressions of the other;
they are also held together by the investing periosteum, or fibrous
membrane, which passes directly from one to the other, and permits
no motion, beyond perhaps a slight yielding to external pressure.
In old animals there is a great tendency for the different bones to
become actually united by the extension of ossification from one to
the other, with consequent obliteration of the sutures. The cranium,
thus formed of numerous originally independent ossifications, which may
retain throughout life more or less of their individuality, or be all
fused together, according to the species, the age, or even individual
peculiarity, consists of a brain-case, or bony capsule for enclosing and
protecting the brain, and a face for the support of the organs of sight,
smell, and taste, and of those concerned in seizing and masticating the
food. The brain-case articulates directly with the anterior cervical
vertebra, by means of a pair of oval eminences, called condyles, placed
on each side of the large median foramen which transmits the spinal cord.
It consists of a basal axis, continuous serially with the axes or centra
of the vertebræ, and of an arch above, roofing over and enclosing the
cavity which contains the cephalic portion of the central nervous system
(see Fig. 6). The base with its arch is composed of three segments placed
one before the other, each of which is comparable to a vertebra with a
greatly expanded neural arch. The hinder or occipital segment consists
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