An introduction to the study of mammals living and extinct — John Shaqi
An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
Although the vertebræ of different regions of the column of the same
animal or of different animals present great diversities of form, yet
there is a certain general resemblance among them, or a common plan on
which they are constructed, which is more or less modified by alteration
of form or proportions, or by the addition or suppression of parts to
fit them to fulfil their special purpose in the economy. An ordinary or
typical vertebra consists, in the first place, of a solid piece of bone,
termed the body or centrum (Fig. 9, _c_), of the form of a disk or short
cylinder. The bodies of contiguous vertebræ are connected together by
a very dense, tough, and elastic material called the “intervertebral
substance,” of peculiar and complex arrangement. This substance forms
the main, and in some cases the only, union between the vertebræ. Its
elasticity provides for the vertebræ always returning to their normal
relation to each other and to the column generally, when they have been
disturbed therefrom by muscular action. A process (_p_) arises on each
side from the dorsal surface of the body. These processes, meeting
in the middle line above, form an arch, surmounting a space or short
canal (_nc_). Since it contains the posterior prolongation of the great
cerebro-spinal nervous axis, or spinal cord, this space is called the
neural canal, and the arch the neural arch, in contradistinction to
another arch on the ventral surface of the body of the vertebræ, called
the hæmal arch. The latter is, however, never formed in mammals by
any part of the vertebra itself, but by certain distinct bones placed
more or less in apposition to it, namely the ribs in the thoracic, and
the “chevron bones” in the caudal region. In most cases the arch of
one vertebra is articulated with that of the next by distinct surfaces
with synovial joints, placed one on each side, called “zygapophyses”
(_az_, _pz_), but these are often entirely wanting when flexibility is
more needed than strength, as in the greater part of the caudal region
of long-tailed animals. In addition to the body and the arch, there
are certain projecting parts called processes, chiefly serving for the
attachment of the numerous muscles which move the vertebral column. Of
these two are single and median, viz. the spinous process, neural spine,
or neurapophysis (_s_), arising from the middle of the upper part of
the arch, and the hypapophysis from the under surface of the body. The
latter, however, is as frequently absent as the former is constant. The
other processes are paired and lateral. They are the transverse processes
(_t_), of which there may be two, an upper and a lower, in which case the
former is called, in the language of Owen (to whom we are indebted for
the terminology of the parts of vertebræ in common use), “diapophysis,”
and the latter “parapophysis.” Other processes less constantly present
are called respectively “metapophyses” (_m_) and “anapophyses” (_a_).
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