The two ridges and that part of the lateral surface of the bone included
between them form a part of the sphenoid bone known as the pterygoid
process of the sphenoid bone, in those cases where the pterygoid is a
separate bone.
The remainder of the process is equivalent to the pterygoid bone of
other vertebrates.
Between the caudal margin of this bone and the lateral of the two
ridges, i.e., between the pterygoid bone and the pterygoid process of
the sphenoid, is a long deep fossa, the =internal pterygoid fossa= (Fig.
40, _s_). The laterocaudal margin of the pterygoid process projects
caudad, as a curved triangular spine, the =hamulus= or =hamular process=
(Fig. 40, _t_; Fig. 43, _i_).
The =Presphenoid Bone=. =Os presphenoidale= (Fig. 21).--In a young cat
this bone is in three pieces, a basal portion (=presphenoid=) and two
wings (=orbitosphenoid bones=). These bones remain distinct throughout
life in many lower vertebrates, but in the adult cat they fuse to form a
single bone. We may nevertheless conveniently describe this bone as made
up of a body (_a_) (the basisphenoid), and two wings (_b_), the
orbitosphenoids (the alæ parvæ of the human sphenoid).
[Illustration: FIG. 21.--PRESPHENOID BONE, VENTRAL VIEW.
_a_, body; _b_, wings; _c_, optic foramina.]
The body (_a_) lies in the base of the skull in the median line, craniad
of the basisphenoid. It has the form of a rectangular prism about twice
as long as broad. It is hollow, and the cavity is divided by a median
longitudinal partition into two cavities (=sphenoidal sinuses=, Fig. 43,
_l_). The sphenoidal sinuses are continued craniad into the cavities of
the ethmoid. The body has six surfaces:
The dorsal or internal surface (Fig. 42, _n_) looks into the cranial
cavity and is continuous with the dorsal surface of the wings. The
caudal end of the body is depressed, and when united to the basisphenoid
aids in forming the cranial wall of the sella turcica. At each
caudolateral angle is a short spine, the =anterior clinoid process=. At
about one-third the length of the bone from the caudal end is a
transverse groove (=chiasmatic groove=, Fig. 42, _m_) for the optic
chiasma. Its ends lead into two round foramina (the =optic foramina=,
Fig. 42, _l_; Fig. 21, _c_) which pass craniolaterad between the body
and the wings of the presphenoid and transmit the optic nerve and the
ophthalmic artery.
The ventral surface (Fig. 21) is hour-glass-shaped and marked by a
smooth median ridge, continuous with the ridge on the basisphenoid and
overlaid at its cranial end by the vomer. The caudal end presents a
rough triangular area on each side, for articulation with the pterygoid
process of the sphenoid bone, while the cranial end has similar areas
overlaid by the nasal portion of the palatine bones.
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