Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
When we turn to the Cordaiteæ we come to a group of plants which bears
distinct relationship to the preceding, but which has a number of
individual characters. It is a group of which we should know nothing were
it not for the fossils preserved in the Palæozoic rocks; yet,
notwithstanding the fact that it flourished so long ago, it is a family
of which we know much. At the time of the Coal Measures and the
succeeding Permo-carboniferous period, it was of great importance, and,
indeed, in some of the French deposits it would seem as though whole
layers of coal were composed entirely of its leaves.
Among the fossil remains of this family there are impressions, casts, and
true petrifactions, so that we know both its external appearance and the
internal anatomy of nearly every part of several species of the genus.
For a long time the various fossil remains of the plant were not
recognized as belonging to each other and together forming the records of
one and the same plant—the broad, long leaves with their parallel veins
were looked on as Monocotyledons (see fig. 61); the pith casts (see fig.
63) were thought to be peculiar constricted stems, and were called
_Sternbergia_; while the wood, which was known from its microscopic
structure, was called _Araucarioxylon_—but the careful work of many
masters of fossil botany, whose laborious studies we cannot describe in
detail here, brought all these fragments together and proved them to
belong to _Cordaites_.
[Illustration: Fig. 61.—Leaf of _Cordaites_, _l_, attached by its broad
base to a Stem, _s_]
We now know that _Cordaites_ were large trees, with strong upright shafts
of wood, to whose branches large simple leaves were attached. The leaves
were much bigger than those of any living Gymnosperm, even than those of
the Kauri Pine (a member of the Araucariaceæ), and seem in some species
to have exceeded 3 ft. in length. The trees branched only at the top of
the main shaft, and with their huge sword-like leaves must have differed
greatly in appearance from any plant now living. The leaves had many
parallel veins, as can be seen in fig. 61, and were attached by a broad
base directly to the main stem; thus coming closer to the Araucarias than
the other groups of Gymnosperms in their leaf characters.
[Illustration: Fig. 62A.—Microscopic Section of Part of a Leaf of
_Cordaites_
V, Vascular bundle; W, wood of bundle; _sh_, its sheath; S^1, large
sclerenchyma mass alternating with bundles; S^2 and S^3, sclerenchyma
caps of bundle; P, soft tissue of leaf.]
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