Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
The internal anatomy is often well preserved, and there is a number of
species of leaves whose anatomy is known. As will be expected from the
parallel veins, in each section there are many vascular bundles running
equidistantly through the tissue. Fig. 62A shows the microscopic details
from a well-preserved leaf. In all the species patches of sclerenchyma
were developed, and everything indicates that they were tough and well
protected against loss of water, even to a greater extent than are most
of the leaves of living Gymnosperms.
In the stems the pith was much larger than that in living Gymnosperms
(where the wood is generally very solid), and it was hollow in older
stems, except for discs of tissue across the cavity. The internal cast
from these stems has been described before, and is seen in fig. 63.
[Illustration: Fig. 62B.—Much-magnified Wood Elements from _Cordaites_
Stem seen in longitudinal section, the type known as _Araucarioxylon_.
Note the hexagonal outlines of the bordered pits, which lie in several
rows]
The wood was formed in closely packed radiating rows by a normal cambium
(see p. 66), and the tracheæ so formed had characteristic rows of
bordered pits (see fig. 62B). The wood comes nearer to that of the living
Araucarias than any other, and indeed the numerous pieces of fossil wood
of this type which are known from all the geological periods are called
_Araucarioxylon_.[11] A double strand goes out from the main mass of
wood, which afterwards divides and subdivides to provide the numerous
bundles of the leaf.
[Illustration: Fig. 63.—Cast of Hollow Pith of _Cordaites_, the
constrictions corresponding to discs of solid tissue across the cavity]
In the case of these fossils we are fortunate enough to have the
fructifications, both male and female, in a good state of preservation.
As in other Gymnosperms, the male and female cones are separate, but they
differed less from each other in their arrangement than do those of any
of the living types hitherto mentioned. They can hardly be described as
true cones, though they had something of that nature; the seeds seem to
be borne on special short stems, round which are also sterile scales. In
the seed and the way it is borne perhaps the Cordaiteæ may be compared
more nearly with the Taxeæ than with the other groups. A seed, not yet
ripe, is shown in slightly diagrammatic form in fig. 64, where the
essential details are illustrated. The seeds of this family sometimes
reached a considerable size, and had a fleshy layer which was thick in
comparison with the stone, and externally comparable with a
cherry—though, of course, of very different nature in reality, for
_Cordaites_, like _Taxus_, is a Gymnosperm, with simple naked seeds,
while a cherry is the fruit of an Angiosperm.
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