In the busy hive of cells which constitutes what we call the animal
body, there is thus ceaseless activity. During periods of apparent
rest the protoplasm is engaged in constructive work, building up fresh
supplies of unstable materials, which, during periods of apparent
activity, break up into simpler and more stable substances, some of
which are useful to the organism, while others must be got rid of as
soon as possible. From another point of view, the cells during apparent
rest are storing up energy to be utilized by the animal during its
periods of activity. The storing up of available energy may be likened
to the winding up of a watch or clock; it is when an organ is at rest
that the cells are winding themselves up; and thus we have the apparent
paradox that the cell is most active and doing most work when the organ
of which it forms a part is at rest. During the repose of an organ, in
fact, the cells are busily working in preparation for the manifestation
of energetic action that is to follow. Just as the brilliant display
of intellectual activity in a great orator is the result of the silent
work of a lifetime, so is the physical manifestation of muscular power
the result of the silent preparatory work of the muscle-cells.
It may, perhaps, seem strange that the products of cellular life should
be reached by the roundabout process of first producing unstable
compounds, from which are then formed more stable substances, useful
for permanent purposes as in bone, or temporary purposes as in the
digestive fluids. It seems a waste of power to build up substances
unnecessarily complex and stored with an unnecessarily abundant supply
of energy. But only thus could the organs be enabled to act under the
influence of stimuli, and afford examples of corporate behaviour. They
are like charged batteries ready to discharge under the influence of
the slightest organic touch. In this way, too, is afforded a means
by which the organ is not dependent only upon the products of the
immediate activity of the protoplasm at the time of action, but can
utilize the store laid up during preceding periods of rest.
Sufficient has now been said to illustrate the nature of some of the
physical processes which accompany organic behaviour in its corporate
aspect. The fact that should stand out clearly is that the animal body
is stored with large quantities of available energy resident in highly
complex and unstable chemical compounds, elaborated by the constituent
cells. These unstable compounds, eminently explosive according to
our analogy, are built up of materials derived from two different
sources--from the nutritive matter (containing carbon, hydrogen, and
nitrogen) absorbed during digestion, and from oxygen taken up from the
air during respiration. The cells thus become charged with energy that
can be set free on the application of the appropriate stimulus, which
may be likened to the spark that fires the explosive.
Public-domain text, read in full here on John Shaqi.
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