[Illustration: FIG. 12.--Solitary Wasp dragging a Caterpillar to its
Nest (after Peckham).]
The mode of carrying their booty is in these wasps instinctive, and
relatively uniform. _Ammophila urnaria_ grasps the caterpillar, near
the anterior end, in her mandibles, and carries or drags it beneath her
legs, walking forwards. It is generally but not always with the ventral
surface uppermost. _Pompilus_ takes hold of her spider anywhere, but
always drags it over the ground, walking backwards. _Oxybelus_ clasps
her fly with her hind legs; _Bembex_ with the second pair. Each works
after her own fashion in a way that is relatively uniform for each
species.
The general style of the nest, its mode of construction, and its method
of closure, are always performed, says Dr. Peckham, by each species
in a similar manner, not indeed in circumstantial detail, but quite
in the same way in a broad sense. Variation or modification is always
present, but the tendency to depart from a nest of a given type is not
excessive. Some dig in the ground curved tunnels, with or without one
or more chambers. Others bore into decaying wood; others use straws,
or make tunnels in bramble stems; while the mud-daubers build cells in
which to store the food and lay the egg. This is sometimes deposited
on the first, sometimes on the last, sometimes on some intermediate
victim, but generally in much the same place and position. _Ammophila_,
for instance, lays it on the side of the sixth or seventh segment--that
is to say, in about the mid position.
Some species first capture their prey, and then make the nest in which
it is to be entombed. Others first prepare the nest, and then carry
or drag their prey to it--often from considerable distances--quite
irrespective of what seems to us the more appropriate method of the
two under the particular circumstances of the case. And the way in
which the victim is dragged into the nest is similarly a matter of
inheritance. Each way is characteristic of the species concerned, and
would be an important part of any definition of the animal based upon
its modes of behaviour. For example, a _Sphex_ places her grasshopper
just at the entrance of the nest, which she then enters herself before
dragging in her prey by the antennæ. When the wasp was in the hole,
Fabre moved the victim a little way off; the wasp came out, brought
the grasshopper to the entrance as before, and went in a second time.
This was repeated about forty times, each time with the same result,
until the patience of the naturalist was exhausted, and the persistent
wasp took her booty in after her appropriate fashion. She must place
the grasshopper close to the opening; she must then descend and examine
the nest, and, after that, must drag it down. Nothing less than the
performance of these acts in a certain order satisfies her instinctive
impulse.
Public-domain text, read in full here on John Shaqi.
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